Speyeria

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Goto:

Speyeria Scudder, 1872

Speyeria Scudder, 1872; 4th Ann. Rep. Peabody Acad. Sci. (1871): 44; TS: Papilio idalia Drury
Semnopsyche Scudder, 1875; Bull. Buffalo Soc. nat. Sci. 2: 238, 258; TS: Papilio diana Cramer
- = Speyeria; [NL4A], 261; Pelham, 2008, J. Res. Lepid. 40: 306; Dunford, 2009, Insecta Mundi 90: 7; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [Wahlberg, 2023]
Subgenus Mesoacidalia Reuss, 1926 (Acidalia); Dt. ent. Zs. 1926 (1): 69; TS: Papilio aglaja Linnaeus
- = Speyeria; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [Wahlberg, 2023]
- = Argynnis; [RFEL]
Subgenus Proacidalia Reuss, 1926 (Acidalia); Dt. ent. Zs. 1926 (1): 69; TS: Argynnis clara Blanchard
- = Speyeria; [Wahlberg, 2023]
Neoacidalia Reuss, 1926; Dt. ent. Zs. 1926 (1): 69; TS: Papilio cybele Fabricius
- = Speyeria; Pelham, 2008, J. Res. Lepid. 40: 306; Dunford, 2009, Insecta Mundi 90: 7

Subgenus Semnopsyche (Acidalia); Reuss, 1926, Dt. ent. Zs. 1926 (1): 69
Subgenus Speyeria (Argynnis); [RFEL]; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 17
Mesoacidalia; [LepIndex]; [CoL] ; [NHM-BMW]
Neoacidalia; [NHM-BMW]
Proacidalia; [NHM-BMW]
Semnopsyche; [NHM-BMW]
Speyeria; [LepIndex]; [CoL] ; [iNaturalist] ; [NHM-BMW]
Speyeria (Argynnina); Pelham, 2008, J. Res. Lepid. 40: 306
Speyeria (Argynnini); [NL4A], 261; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [Wahlberg, 2023]

700x734 (~72Kb) USA: nr Crystal Mountain (47°13'26N 120°40'11W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela

600x503 (~56Kb) USA: Cape Blanco, (42°49'57N 124°32'50W ±1km), Curry Co. OR, 2.8.2001, Photo © Markku Savela

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.

Speyeria aglaja (Linnaeus, 1758)

EU, Morocco, Iran - Siberia, AC, China, Korea, Japan. See [maps] Dostojka aglaja Le Grand Nacré Orvokkihopeatäplä Dukatpärlefjäril Großer Perlmutterfalter Stor pärlemorfjäril Dark Green Fritillary Lunares de Piata

Papilio aglaja Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 481; TL: Sweden
Papilio pasithoe Linnaeus, 1767; Syst. Nat. (Edn 12) 1 (2): 755 (repl. for Papilio aglaja Linnaeus)
Papilio emilia Acerbi, 1802; Travels North Cape 2: 253, pl. 2, f. 1-2
Papilio charlotta Haworth, 1802; Prodr. Lep. Brit.: 3
jurassina (Ragonot, 1871); Rev neuchateloise: [?]
Argynnis aglaja ab. eridioides Pflümer, 1879; Stettin Ent. Ztg 40: 157
Argynnis aglaja ab. aberrans Lampa, 1885; Ent. Tidskr. 6 (1): 20; TL: Sweden, Jmt., Åreskutan
Argynnis aglaja ab. wimani Holmgren, 1888; Ent. Tidskr. 9: 103, f. 1
arvernensis Bramson, 1890²; Tagfalter Eur. Caucasus: 72
Argynnis aglaja ab. suffusa Tutt, 1896; Brit. Butts: 290
Argynnis aglaja ♀ ab. aurea Tutt, 1896; Brit. Butts: 291
Argynnis aglaja ab. flavescens Tutt, 1896; Brit. Butts: 291
Argynnis aglaja ab. pallida Tutt, 1896; Brit. Butts: 291
Argynnis aglaja ab. fusca Tutt, 1896; Brit. Butts: 292
Argynnis aglaja methana Fruhstorfer, 1908; Ent. Zs. 22 (39): 161; TL: Sierra de Guadarrama, Castilia
Argynnis aglaia v. basalis Matsumura, 1908; Ent. Zs. 22 (39): 158; TL: Hokkaido
Argynnis aglaia ab. albomaculata Rebel, [1909]; in Berge, Schmetterlingsbuch (edn 9): 33
Argynnis aglaja plutus Oberthür, 1909; Étud. Lépid. Comp. 3: 209 (Leech, 1983 pl. 22, f. 6 ♀); TL: W.China, Wa-ssu-kow
Argynnis aglaja ab. hortensia Riel, 1911; Ann. Soc. Linn. Lyon (n.s.) 58: 187 [ Donzel?]
Argynnis aglaia ab. gutta Wileman, 1911; Entomologist 44: 55; TL: Yezo, Jozanke
Argynnis aglaia ashretha Evans, 1912; J. Bombay nat. Hist. Soc. 21 (2): 558, 582; TL: Chitral, Ashreth Valley
Argynnis aglaja ab. viridiatra Strand, 1912; Ent. Zs. 25 (49): 253
Argynnis aglaja yopala Fruhstorfer, 1912; in Seitz, Gross-Schmett. Erde 9: 515; TL: Chitral
Argynnis aglaja taldena Fruhstorfer, 1912; in Seitz, Gross-Schmett. Erde 9: 516; TL: Setchuan
?Argynnis aglaja appenninicola Verity, 1914; Boll. Soc. ent. ital. 45: 213, f. 4-5
Argynnis aglaia ab. molybdina Newnham, 1917; Entomologist 50: 230
Argynnis aglaja r. locuples Verity, 1919; Ent. Rec. J. Var. 31: 195
Argynnis aglaja r. emilocuples Verity, 1919; Ent. Rec. J. Var. 31: 195
Argynnis aglaia r. locupletata Verity, 1922; Ent. Rec. J. Var. 34: 15 (repl. Argynnys aglaja r. locuples Verity, 1919)
scotica (Watkins, 1923) (Argynnis); Entomologist 56: 105
Argynnis aglaia v. purpura Evans, 1924; J. Bombay nat. Hist. Soc. 30 (1): 89
Argynnis aglaja mirabilis de Sagarra, 1925; Butll. Inst. catal. Hist. nat. (2) 5 (1): 45; TL: Orihuela del Tremedal (Aragó)
Argynnis aglaja montesignum de Sagarra, 1926; Butll. Inst. catal. Hist. nat. (2) 6 (6-7): 132
valesinoides (Reuss, 1926) (Acidalia); Dt. ent. Zs. 1926 (1): 69; TL: Kashmir
Argynnis aglaja ab. hindenburgi Schuster von Forstner, 1928; Lepid. Rdsch. 2 (16): 157
ovalis (Cabeau, 1930) (Argynnis); Lambillionea 30: 179
Argynnis aglaja f. clorinda de Sagarra, 1930; Butll. Inst. catal. Hist. nat. (2) 10 (7): 115
Argynnis aglaja r. fuscans Verity, 1935; Entomologist 68: 190; TL: Alpi Carniche, Sappada, 1300m
Argynnis aglaja r. rubidior Verity, 1935; Entomologist 68: 190; TL: Lago Maggiore
Argynnis aglaja r. subvitatha Verity, 1935; Entomologist 68: 202; TL: Tianchan, Aksultal
linnaei (Hemming, 1942) (Mesoacidalia); Proc. R. ent. Soc. Lond. (B) 11 (11): 159
Argynnis aglaja f. ocellata Eisner, 1942; Zool. Meded. Leiden 24 (4): 123; TL: Krummhübel, Riesengebirge
Argynnis aglaja f. duplicata Eisner, 1942; Zool. Meded. Leiden 24 (4): 123; TL: Amata, Lettland
Argynnis aglaja kansuensis Eisner, 1942; Zool. Meded. Leiden 24 (4): 123; TL: Sining, Kansu, 2500m
Argynnis aglaja ab. smrzi Slaby, 1949; Acta Soc. ent. Cal. 45 (1-2): 68
Mesoacidalia charlotta f. pluriradiata Verity, 1950; Le Farfalle diurn. d'Italia 4: 301
?rubescens (Verity, 1950) (Mesoacidalia); Le Farfalle diurn. d'Italia 4: 302
Argynnis aglaia ab. robnora Kershaw, 1952; Ent. Rec. J. Var. 64: 69, pl. 3, f. 15
Argynnis aglaia ab. mitchelli Kershaw, 1952; Ent. Rec. J. Var. 64: 69, pl. 3, f. 16
Mesoacidalia charlotta f. cadmeis Lempke, 1956; Tijdschr. Ent. 99 (4): 175
Mesoacidalia charlotta f. transversa Lempke, 1956; Tijdschr. Ent. 99 (4): 175
Mesoacidalia charlotta f. infraochracea Lempke, 1956; Tijdschr. Ent. 99 (4): 175
sinenigra (Aagesen, 1977) (Mesoacidalia); Lepidoptera Kbh. 3 (2-3): 48

Argynnis (Speyeria) aglaja; [RFEL]
Argynnis aglaja methana; [iNaturalist]
Argynnis aglaja ocellata; [iNaturalist]
Argynnis aglaja rubidior; [iNaturalist]
Argynnis aglaja; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 439; [SPRK]; [BRU2]: 36, pl. 23, f. 1-15; [Otakar Kudrna]; Yakovlev, 2012, Nota lepid. 35 (1): 92; [iNaturalist] ; [Mario Maier]
Argynnis aglaja ab. wimani; Fruhstorfer, 1906, Ent. Zs. 20 (3): 17
Mesoacidalia aglaia[sic, aglaja]; [EBW]
Mesoacidalia aglaja; [H&R], 91; [BOW]: pl. 3, f. 13-14; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
Mesoacidalia vitatha ashretha; [LepIndex]; [CoL]
Speyeria aglaia basalis; [LepIndex]
Speyeria aglaia cadmeis; [LepIndex]
Speyeria aglaia clorinda; [LepIndex]
Speyeria aglaia duplicata; [LepIndex]
Speyeria aglaia fuscans; [LepIndex]
Speyeria aglaia hortensia; [LepIndex]
Speyeria aglaia infraochracea; [LepIndex]
Speyeria aglaia jurassina; [LepIndex]
Speyeria aglaia kansuensis; [LepIndex]
Speyeria aglaia linnaei; [LepIndex]
Speyeria aglaia locupletata; [LepIndex]
Speyeria aglaia methana; [LepIndex]
Speyeria aglaia mirabilis; [LepIndex]
Speyeria aglaia montesignum; [LepIndex]
Speyeria aglaia ocellata; [LepIndex]
Speyeria aglaia pluriradiata; [LepIndex]
Speyeria aglaia purpura; [LepIndex]
Speyeria aglaia rubidior; [LepIndex]
Speyeria aglaia scotica; [LepIndex]
Speyeria aglaia subvitatha; [LepIndex]
Speyeria aglaia taldena; [LepIndex]
Speyeria aglaia transversa; [LepIndex]
Speyeria aglaia yopala; [LepIndex]
Speyeria aglaja appennicola; [CoL]
Speyeria aglaja basalis; [CoL]
Speyeria aglaja cadmeis; [CoL]
Speyeria aglaja clorinda; [CoL]
Speyeria aglaja duplicata; [CoL]
Speyeria aglaja fuscans; [CoL]
Speyeria aglaja hortensia; [CoL]
Speyeria aglaja infraochracea; [CoL]
Speyeria aglaja jurassina; [CoL]
Speyeria aglaja kansuensis; [CoL]
Speyeria aglaja linnaei; [CoL]
Speyeria aglaja locupletata; [CoL]
Speyeria aglaja methana; [CoL]
Speyeria aglaja mirabilis; [CoL]
Speyeria aglaja montesignum; [CoL]
Speyeria aglaja ocellata; [CoL]
Speyeria aglaja pluriradiata; [CoL]
Speyeria aglaja purpura; [CoL]
Speyeria aglaja rubidior; [CoL]
Speyeria aglaja scotica; [CoL]
Speyeria aglaja semilocuples; [CoL]
Speyeria aglaja subvitatha; [CoL]
Speyeria aglaja taldena; [CoL]
Speyeria aglaja transversa; [CoL]
Speyeria aglaja yopala; [CoL]
Speyeria aglaja; [MRS], 473; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [CoL] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [SVF]

500x363 (~61Kb) upperside West down, Devo, UK 14/06/95, Photo © Simon Coombes	500x389 (~50Kb) underside West down, Devo, UK 14/06/95, Photo © Simon Coombes	700x544 (~41Kb) upperside Finland, Kangasala, Ruutana, 682:34, 8/1997, Photo © Tero Piirainen
780x696 (~49Kb) FINLAND: Ab: Suomusjärvi, 669:31, 29.6.1997, Photo © Markku Savela	500x667 (~41Kb) ova Finland, Sipoo 670:39, 22.7.1980, Photo © Pertti Pakkanen	550x371 (~71Kb) larva Central Europe 1990-2000, Photo © Michael Borsch
550x371 (~80Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch	597x400 (~59Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch	777x565 (~100Kb) ♀ upperside Russia, Moscow area, 29.7.2008, Photo © D. Smirnov
551x607 (~112Kb) ♀ upperside Russia, Moscow area, 29.7.2008, Photo © D. Smirnov

Larva on Viola , V. canina, V. riviniana, V. tricolor [SPRK], Viola bicolor, Bistorta major [BRU2]: 36, Viola verecunda, Polygonum suffultum [MRS]

S. a. aglaja SEU, CEU, Altai, Sayan, W.Siberia, S.Siberia
- Argynnnis aglaja aglaja; [BRU2], 36
- Mesoacidalia aglaja aglaja; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
S. a. vitatha (Moore, [1875]) Pamirs
- Argynnis vitatha Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4): 568; TL: Cashmere
- ?Argynnis aglaja cannelata Peschke, 1934; Int. ent. Zs. 28: 431; TL: Kashmir
- Argynnis aglaja vitatha; [BRU2], 36
- Mesoacidalia aglaja vitatha; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
- Mesoacidalia vitatha; [EBW], (name only); [LepIndex] ; [CoL]
- Speyeria aglaja vitatha; [MRS], 473
S. a. fortuna (Janson, 1877) Japan
- Argynnis fortuna Janson, 1877; Cistula ent. 2: 154; TL: Japan
- Argynnis aglaja fortuna f. myonia Fruhstorfer, 1907; Int. ent. Zs. 1 (34): 257; TL: Nagasaki
- Argynnis aglaia fortuna ab. shanae Kuwayama, 1921; Dobuts. Z. 33: 333, 336, f. 3; TL: Etoroff I.
- Argynnis aglaia fortuna ab. wilemana Kuwayama, 1921; Dobuts. Z. 33: 334, 336, f. 4; TL: Sapporo
- Argynnis aglaia myonia ab. isshikii Matsumura, 1928; Insecta Matsumurana 2 (4): 193; TL: Hokkaido
- Mesoacidalia aglaja fortuna; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
- Mesoacidalia charlotta fortuna; Kudrna, 1974, Atalanta 5: 107
- Speyeria aglaia fortuna; [LepIndex]
- Speyeria aglaja fortuna; [CoL]
S. a. ottomana (Röber, 1896) Armenia, Talys, Kopet-Dagh
- Argynnis aglaja var. ottomana Röber, 1896; Ent. Nachr. 22 (6): 82
- Argynnis aglaja ottomana; [BRU2], 36; [iNaturalist]
- Speyeria aglaia ottomana; [LepIndex]
- Speyeria aglaja ottomana; [CoL]
S. a. kenteana (Stichel, 1901) Transbaikalia, N.Ussuri, Amur
- Argynnis aglaja kenteana Stichel, 1901; Berl. ent. Z. 46: 23 (Stgr. i. l.)
- Argynnis aglaja graeseri Kardakoff, 1928; Ent. Mitt. 17 (4): 270, pl. 7, f. 7-8; TL: Ussuri
- Argynnis aglaja kenteana; [BRU2], 36; [RFEL]
- Mesoacidalia aglaja kenteana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
- Speyeria aglaia graeseri; [LepIndex]
- Speyeria aglaja graeseri; [CoL]
S. a. bessa (Fruhstorfer, 1907)
- Argynnis aglaja bessa Fruhstorfer, 1907; Int. ent. Zs. 1 (34): 257; TL: Tse-Kou
- Argynnis aglaja bessa; [iNaturalist]
- Speyeria aglaia bessa; [LepIndex]
- Speyeria aglaja bessa; Huang, 2001; [CoL]
S. a. auxo (Jachontov, 1909) Caucasus, Transcaucasus
- Argynnis aglaja var. auxo Jachontov, 1909; Revue Russe Ent. 8: 287; TL: Caucasus
- Mesoacidalia aglaja auxo; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
- Speyeria aglaia auxo; [LepIndex]
- Speyeria aglaja auxo; [CoL]
S. a. lyauteyi (Oberthür, 1920) Morocco (Middle Atlas)
- Argynnis lyauteyi Oberthür, 1920; Étud. Lépid. Comp. 17: 48, pl. D; TL: Azrou (Morocco)
- Mesoacidalia aglaja lyauteyi; [BMAT]: 50, pl. 15, f. 10-13
- Mesoacidalia liauteyi[sic, lyauteyi]; [EBW], (name only)
- Speyeria aglaia lyauteyi; [LepIndex]
- Speyeria aglaja lyauteyi; [CoL]
435x600 (~32Kb) upperside Morocco, Ifrane, Moyen Atlas, Photo © Michel Tarrier 600x435 (~33Kb) underside Morocco, Ifrane, Moyen Atlas, Photo © Michel Tarrier
S. a. matsumurai (Nakahara, 1926) Kuriles
- Argynnis aglaja matsumurai Nakahara, 1926; Ins. Inscit. Mens. 14: 48
- Argynnis aglaia chishimensis Matsumura, 1928; Insecta Matsumurana 2 (4): 192
- Argynnis aglaja matsumurai; [BRU2], 36
- Speyeria aglaia chishimensis; [LepIndex]
- Speyeria aglaia matsumurai; [LepIndex]
- Speyeria aglaja chishimensis; [CoL]
- Speyeria aglaja matsumurai; [CoL]
S. a. tonnai (Matsumura, 1928) Sakhalin
- Argynnis aglaia var. sachalinensis Matsumura, 1911; J. Coll. Agr. Tohoku Imp. Univ. Sapporo 4 (1): 43 (preocc.)
- Argynnis aglaia[sic] tonnai ab. otaniana Matsumura, 1928; Insecta Matsumurana 2 (4): 192; TL: Saghalien
- Argynnis aglaja tonnai; [BRU2], 36
- Speyeria aglaja otaniana; [CoL]
S. a. clavimacula (Matsumura, 1929) S.Ussuri
- Argynnis aglaia[sic] clavimacula Matsumura, 1929; Insecta Matsumurana 3 (4): 154; TL: Korea
- Argynnis aglaja clavimacula; [BRU2], 36
- Speyeria aglaia clavimacula; [LepIndex]
- Speyeria aglaja clavimacula; [MRS], 473; [CoL]
S. a. excelsior (Rothschild, 1933) Morocco (Rif Mts.)
- Argynnis aglaia excelsior Rothschild, 1933; Novit. Zool. 38: 317; TL: Ketama (Morocco)
- Argynnis lyauteyi excelsior;
- Mesoacidalia aglaja excelsior; [BMAT]: 50, pl. 15, f. 14-17
- Speyeria aglaia excelsior; [LepIndex]
- Speyeria aglaja excelsior; [CoL]
600x435 (~46Kb) upperside Morocco, Djebel Tisirène, Rif, Photo © Michel Tarrier 600x435 (~29Kb) underside Morocco, Djebel Tisirène, Rif, Photo © Michel Tarrier
S. a. gigasvitatha (Verity, 1935) Tian-Shan, Ghissar, Darvaz, Alai, S.Altai
- Argynnis aglaja gigasvitatha Verity, 1935; Entomologist 68: 202; TL: Issyk kul
- Argynnis aglaja alaicola Verity, 1935; Entomologist 68: 202
- Argynnis aglaja gigasvitatha; [BRU2], 36
- Speyeria aglaia alaicola; [LepIndex]
- Speyeria aglaia gigasvitatha; [LepIndex]
- Speyeria aglaja alaicola; [CoL]
- Speyeria aglaja gigasvitatha; [CoL]
S. a. boreas (Hemming, 1942) EU, Siberia, Far East, Kamchatka
- Argynnis aglaja v. borealis Strand, 1901; Nyt Mag. Naturv. 39 (1): 49 (preocc. Argynnis thore boralis Staudinger, 1861)
- boreas (Hemming, 1942) (Mesocacidalia); Proc. R. ent. Soc. Lond. (B) 11 (11): 160 (repl. Argynnis aglaja borealis Strand, 1901)
- Argynnis aglaja borealis; [BRU2], 36; [RFEL]; [iNaturalist]
- Mesocacidalia aglaja boreas; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
- Speyeria aglaia boreas; [LepIndex]
- Speyeria aglaja boreas; [CoL]

Speyeria alexandra (Ménétriés, 1832) N.Iran, W.Kopet-Dagh, Asia Minor. See [maps]
- Argynnis alexandra Ménétriés, 1832; Cat. Rais.: 246; TL: [Zuvand Plateau, Talysh Mts., Azerbaijan]
- Argynnis alexandra; [BRU2]: 36, pl. 22, f. 7-9; [iNaturalist]
- Fabriciana alexandra; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
- Mesoacidalia alexandra; [BOW]: pl. 198, f. 5
- Speyeria aglaia alexandra; [LepIndex]
- Speyeria aglaja alexandra; [CoL]
- Speyeria alexandra; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LEPIFORUM]
Speyeria clara (Blanchard, [1844]) -. See [maps]
- Argynnis clara Blanchard, [1844]; in Jacquemont, Voy. Inde 4 (Zool.): 20, pl. 2, f. 2-3
- Argynnis clara; de Nicéville, 1889, J. Bombay nat. Hist. Soc. 4 (3): 165, pl. A, f. 6 ♀; [iNaturalist]
- Mesoacidalia clara; [LepIndex] ; [CoL]
- Speyeria clara; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643
- S. c. clara Kashmir, NW.India
  - Mesoacidalia clara clara; Lang & Han, 2009, Atalanta 40 (3/4): 494
- S. c. clarina (Staudinger, 1901) E.Qinghai, SW.Gansu, W.Sichuna, E.Tibet
  - Argynnis clara v. clarina Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 38; TL: Amdo
  - Mesoacidalia clara neoclara Chou, Yuan, Yin, Zhang & Chen, 2002; Entomotaxonomia 24 (1): (52-68); TL: Gansu
    
    = Mesoacidalia clara clarina; Lang & Han, 2009, Atalanta 40 (3/4): 494
  - Mesoacidalia clara clarina; [LepIndex]; [CoL]
- S. c. manis (Fruhstorfer, 1903) Sikkim, S.Tibet
  - Argynnis clara manis Fruhstorfer, 1903; Dt. ent. Z. Iris 16 (1): 41; TL: Sikkim
  - Argynnis claudia Fawcett, 1904; Proc. zool. Soc. Lond. 2 (1): 136; TL: Khamba Jong, Thibet
    
    = Mesoacidalia clara manis; Lang & Han, 2009, Atalanta 40 (3/4): 494
  - Argynnis claudia; [LepIndex] ; [CoL] ; [iNaturalist]
  - Mesoacidalia clara manis; Lang & Han, 2009, Atalanta 40 (3/4): 494; [LepIndex]; [CoL]
  - Mesoacidalia claudia; [BOW]: pl. 196, f. 10
- S. c. menba (Lang, 2009) Tibet
  - Mesoacidalia clara menba Lang, 2009; Atalanta 40 (3/4): 494; TL: Tibet, Milin Co., Namula Pass, 3800m
- S. c. tongtianensis (Lang, 2009) Qinghai
  - Mesocacidalia clara tongtianensis Lang, 2009; Atalanta 40 (3/4): 495; TL: Qinghai, Qumalai Co., Chuhuier River, 4500m
- S. c. kanga (Lang, 2010) Sichuan
  - Mesocacidalia clara kanga Lang, 2010; Atalanta 41: 221; TL: Sichuan, Kangding, Muge Cuo Lake

Speyeria diana (Cramer, [1777])

SE.USA. See [maps] Diana

Papilio diana Cramer, [1777]; Uitl. Kapellen 2 (9-16): 4, 148, pl. 98, f. D, E

Semnopsyche diana; Dyar, 1903, Bull. U.S. nat. Mus. 52: 12
Speyeria diana; [NACL], #4449; [NAB], 544; [EBW]; [BOW]: pl. 15, f. 15-16; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 307; Dunford, 2009, Insecta Mundi 90: 11; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [LEPIFORUM] ; [BAMONA]

940x735 (~121Kb) USA: Alabama, 4.6.2005, Photo © Vitaly Charny	1078x876 (~155Kb) USA: Alabama, 4.6.2005, Photo © Vitaly Charny	1203x883 (~192Kb) upperside USA: Alabama, 4.6.2005, Photo © Vitaly Charny
1321x1137 (~214Kb) underside USA: Alabama, 4.6.2005, Photo © Vitaly Charny	1267x989 (~137Kb) USA: Alabama, 4.6.2005, Photo © Vitaly Charny

Larva on Viola papilionacea, V. cucullata, V. cornuta, V. sororia Dunford, 2009, Insecta Mundi 90: 12

Speyeria cybele (Fabricius, 1775)

USA. See [maps] Great Spangled Fritillary

Papilio cybele Fabricius, 1775; Syst. Ent.: 516

Argynnis cybele; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13; [MPG] ; [iNaturalist] ; [BOA]
Speyeria cybele; [NACL], #4450; [NAB], 545; [EBW]; [BOW]: pl. 15, f. 14, 17; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 307; Dunford, 2009, Insecta Mundi 90: 13; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [LEPIFORUM] ; [BAMONA]

1024x761 (~81Kb) upperside USA: Taneum Creek, Wenatchee National Forest, (47°06'24N 120°54'44W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela	700x632 (~45Kb) underside USA: Taneum Creek, Wenatchee National Forest, (47°06'24N 120°54'44W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela	1000x790 (~91Kb) ♂ upperside USA: nr Crystal Mountain / FR9726 (47°14'35N 120°40'41W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela
800x639 (~62Kb) ♂ underside USA: nr Crystal Mountain / FR9726 (47°14'35N 120°40'41W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela	700x986 (~93Kb) ♀ USA: Mima Mounds Natural Area Preserve, Thurston Co., WA, 13.8.2001, Photo © Markku Savela	1300x909 (~199Kb) upperside USA: Alabama, 26.5.2001, Photo © Vitaly Charny
1300x1065 (~179Kb) underside USA: Alabama, 26.5.2001, Photo © Vitaly Charny

Larva on Viola rotundifolia, V. papilionacea, V. palustris, V. adunca, Viola adunca var. bellidifolia, V. sororia, V. canadensis Dunford, 2009, Insecta Mundi 90: 14

S. c. cybele
- Papilio daphnis Cramer, [1775]; Uitl. Kapellen 1 (1-7): 89, pl. 57, f. E, F (preocc. Papilio daphnis [Schiffermüller], 1775)
  - = ; Pelham, 2008, J. Res. Lepid. 40: 307
  - = Speyeria cybele; Dunford, 2009, Insecta Mundi 90: 13
- Argynnis cybele ab. ♂ baal Strecker, 1878; Synon. Cat. Macroplep.: 111
  - = ; Pelham, 2008, J. Res. Lepid. 40: 307
  - = Speyeria cybele; Dunford, 2009, Insecta Mundi 90: 13
- Speyeria cybele cybele; Pelham, 2008, J. Res. Lepid. 40: 307
S. c. leto (Behr, 1862) California, Alberta, Wyoming, ...
- Argynnis leto Behr, 1862; Proc. Cal. Acad. Sci. 2: 173; TL: California
- Argynnis leto var. letis Wright, 1905; Butts. West Coast US: 52; TL: Northern California
  - = ; Pelham, 2008, J. Res. Lepid. 40: 308
- Acidalia (Semnopsyche) valesinoides-alba [= valesinoidesalba] Reuss, 1926; Dt. ent. Zs. 1926 (1): 69
  - = ?; Pelham, 2008, J. Res. Lepid. 40: 308
  - = ?Speyeria nokomis nokomis; Pelham, 2008, J. Res. Lepid. 40: 310
  - = Speyeria nokomis; Dunford, 2009, Insecta Mundi 90: 15
- Argynnis leto tr.f. lethe Gunder, 1934; Can. Ent. 66 (6): 125; TL: Big Horn Mts, Wyoming
- Argynnis leto; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13; [MPG] ; [BOA]
- Speyeria cybele leto; [NACL], #4450h; Pelham, 2008, J. Res. Lepid. 40: 307
- Speyeria leto lethe; [LepIndex]; [CoL]
- Speyeria leto letis; [LepIndex]; [CoL]
- Speyeria leto valesinoidesalba; [LepIndex]; [CoL]
- Speyeria leto; [LepIndex] ; [CoL]
- Speyeria nokomis valesinoalba; [LepIndex]; [CoL]
S. c. carpenterii (Edwards, 1876) New Mexico
- Argynnis carpenterii Edwards, 1876; Trans. amer. ent. Soc. 5: 204; TL: New Mexico
- Argynnis cybele carpenterii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13; [iNaturalist] ; [BOA]
- Speyeria cybele carpenterii; [NACL], #4450d; Pelham, 2008, J. Res. Lepid. 40: 307; [LepIndex]; [CoL]
S. c. charlottii (Barnes, 1897) Colorado
- Argynnis charlottii Barnes, 1897; Can. Ent. 29 (2): 39; TL: Glenwood Spring, Colorado
- Speyeria cybele charlottii; [NACL], #4450e; Pelham, 2008, J. Res. Lepid. 40: 307; [LepIndex]; [CoL]
S. c. krautwurmi (Holland, 1931) Michigan, ...
- Argynnis cybele var. krautwurmi Holland, 1931; Ann. Carnegie Mus. 20 (2): 255; TL: Les Cheneaux, Upper Peninsula of Michigan
- Argynnis cybele krautwurmi; [iNaturalist] ; [BOA]
- Speyeria cybele krautwurmi; [NACL], #4450b; Pelham, 2008, J. Res. Lepid. 40: 307; [LepIndex]; [CoL]
S. c. novascotiae (McDunnough, 1935) Nova Scotia
- Argynnis cybele novascotiae McDunnough, 1935; Can. ent. 67 (1): 18; TL: White Point Beach, Queens Co., Nova Scotia
- Argynnis cybele novascotiae; [iNaturalist] ; [BOA]
- Speyeria cybele novascotiae; [NACL], #4450a; Pelham, 2008, J. Res. Lepid. 40: 307; [LepIndex]; [CoL]
S. c. pseudocarpenteri (Chermock & Chermock, 1940) Manitoba, ...
- Argynnis cybele pseudocarpenteri Chermock & Chermock, 1940; Can. Ent. 72 (4): 82; TL: Sand Ridge, Manitoba
- Argynnis cybele pseudocarpenteri; [iNaturalist] ; [BOA]
- Speyeria cybele pseudocarpenteri; [NACL], #4450c; Pelham, 2008, J. Res. Lepid. 40: 307; [LepIndex]; [CoL]
S. c. letona dos Passos & Grey, 1945 Utah
- Speyeria cybele letona dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 1; TL: City Creek Canyon, Salt Lake City, Utah
- Speyeria cybele letona; [NACL], #4450f; Pelham, 2008, J. Res. Lepid. 40: 307
- Speyeria leto letona; [LepIndex]; [CoL]
S. c. pugetensis Chermock & Frechin, 1947 Washington
- Speyeria cybele pugetensis Chermock & Frechin, 1947; Pan-Pacific Ent. 23 (3): 111; TL: Stimson Creek, nr Belfair, Mason Co., Washington
- Speyeria cybele pugetensis; [NACL], #4450g; Pelham, 2008, J. Res. Lepid. 40: 308
- Speyeria leto pugetensis; [LepIndex]; [CoL]
S. c. eileenae Emmel, Emmel & Mattoon, 1998 California
- Speyeria cybele eileenae Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 144; TL: California, Humboldt Co
- Speyeria cybele eileenae; Pelham, 2008, J. Res. Lepid. 40: 308

Speyeria aphrodite (Fabricius, 1787)

Canada - Kansas. See [maps] Aphrodite Fritillary

Papilio aphrodite Fabricius, 1787; Mantissa Insectorum 2: 62
?daphnis Martyn
- = ; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13

Argynnis aphrodite; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13; [MPG] ; [iNaturalist] ; [BOA] ; [BOA]
Speyeria aphrodite; [NACL], #4451; [NAB], 546; [BOW]: pl. 15, f. 5-6; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 308; Dunford, 2009, Insecta Mundi 90: 14; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [BAMONA]

1000x1262 (~165Kb) underside USA: Pike National Forest, Sugar Creek on CR-67 (about 39°18'N 105°10'W), Douglas Co., CO, 29.7.2012, Photo © Markku Savela

1000x1358 (~194Kb) upperside USA: Pike National Forest, Sugar Creek on CR-67 (about 39°18'N 105°10'W), Douglas Co., CO, 29.7.2012, Photo © Markku Savela

Larva on Viola lanceolata, V. fimbriatula, V. nuttallii, V. papilionacea, V. nephrophylla, Viola primulifolia var. acuta, Viola sagittata, V. sororia, V. tricolor, V. adunca Dunford, 2009, Insecta Mundi 90: 15

S. a. aphrodite New York, Massachusetts, Ohio, ...
- Argynnis cybele f. bartschi Reiff, 1910; Psyche 17 (6): 252; TL: in West Roxbury, Mass.
  - = ; Pelham, 2008, J. Res. Lepid. 40: 308
  - = Speyeria aphrodite; Dunford, 2009, Insecta Mundi 90: 14
- Argynnis aphrodite ab. bakeri Clark, 1932; Bull. U.S. nat. Mus. 157: 276, pl. 19, f. 1-2; TL: Waynesburg, Ohio
  - = ; Pelham, 2008, J. Res. Lepid. 40: 308
  - = Speyeria aphrodite; Dunford, 2009, Insecta Mundi 90: 14
- Speyeria aphrodite ab. hughi Clark & Clark, 1951; Smiths. Misc. Coll. 116 (7): 60
  - = ; Pelham, 2008, J. Res. Lepid. 40: 308
- Speyeria aphrodite aphrodite; Pelham, 2008, J. Res. Lepid. 40: 308
S. a. alcestis (Edwards, [1877]) Illinois, Iow, Colorado, Michigan
- Argynnis alcestis Edwards, [1877]; Trans. amer. ent. Soc. 5: 289
- Argynnis alcestis ab. suffusa Wolcott, 1916; Ent. News, 27 (3): 97; TL: bog S of Grand Rapids, Michigan
  - = ; Pelham, 2008, J. Res. Lepid. 40: 308
- Argynnis aphrodite alcestis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13; [iNaturalist] ; [BOA]
- Speyeria aphrodite alcestis; [NACL], #4451b; Pelham, 2008, J. Res. Lepid. 40: 308; [LepIndex]; [CoL]
S. a. columbia (H. Edwards, 1877) N.British Columbia, ...
- Argynnis columbia H. Edwards, 1877; Proc. Cal. Acad. Sc.: [1]; TL: Lahache, nr Alaskan border of British Columbia
- Argynnis columbia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria aphrodite colombia; [LepIndex]; [CoL]
- Speyeria aphrodite columbia; [NACL], #4451e; Pelham, 2008, J. Res. Lepid. 40: 309
S. a. byblis (Barnes & Benjamin, 1926) Arizona, ...
- Dryas aphrodite r. byblis Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 91; TL: White Mts., Arizona
- Speyeria aphrodite byblis; [NACL], #4451g; Pelham, 2008, J. Res. Lepid. 40: 309; [LepIndex]; [CoL]
S. a. winni (Gunder, 1932) Quebec
- Argynnis aphrodite r. winni Gunder, 1932; Can. Ent. 64 (12): 278; TL: St. Calixte de Kilkenny, Quebec
- Argynnis aphrodite winni; [iNaturalist] ; [BOA]
- Speyeria aphrodite winni; [NACL], #4451a; Pelham, 2008, J. Res. Lepid. 40: 309; [LepIndex]; [CoL]
S. a. whitehousei (Gunder, 1932) British Columbia
- Argynnis whitehousei Gunder, 1932; Can. Ent. 64 (12): 279; TL: Jafrray, B.C.
- Speyeria aphrodite whitehousei; [NACL], #4451d; Pelham, 2008, J. Res. Lepid. 40: 309; [LepIndex]; [CoL]
S. a. ethne (Hemming, 1933)
- Argynnis ethne Hemming, 1933; Entomologist 66 (12): 276 (repl. Argynnis cypris Edwards, 1886)
- Argynnis cypris Edwards, 1886; Can. Ent. 18 (4): 62 (preocc. Argynnis cypris Meigen, 1828)
  - = ; Pelham, 2008, J. Res. Lepid. 40: 309
- Speyeria aphrodite ethne; [NACL], #4451f; Pelham, 2008, J. Res. Lepid. 40: 309; [LepIndex]; [CoL]
S. a. manitoba (Chermock & Chermock, 1940) Manitoba, ...
- Argynnis manitoba Chermock & Chermock, 1940; Can. Ent. 72 (4): 83; TL: Sand Ridge, Manitoba
- Argynnis aphrodite cypris tr.f. mayae Gunder, 1932; Can. Ent. 64 (12): 279; TL: Sand Ridge, Bener Dam Lake, nr Kelwood, Manitoba
  - = ; Pelham, 2008, J. Res. Lepid. 40: 309
- Speyeria (Semnopsyche) aphrodite mayae dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 8; TL: Sand Ridge, Bener Dam Lake, nr Kelwood, Manitoba
  - = ; Pelham, 2008, J. Res. Lepid. 40: 309
- Speyeria aphrodite manitoba; [NACL], #4451c; Pelham, 2008, J. Res. Lepid. 40: 309
- Speyeria aphrodite mayae; [LepIndex]; [CoL]
S. a. cullasaja Gatrelle, 1998 North Carolina, ...
- Speyeria aphrodite cullasaja Gatrelle, 1998; Taxonomic Rep. 1 (3): 2; TL: trail to Scaly Mtn, 4200', Macon Co., North Carolina
- Speyeria aphrodite cullasaja; Pelham, 2008, J. Res. Lepid. 40: 308

Speyeria idalia (Drury, [1773]) E.USA. See [maps] Regal Fritillary
- Papilio idalia Drury, [1773]; Illust. Nat. Hist. Exot. Insects 1: index, 25, pl. 13, f. 1-3
- Speyeria idalia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13; [NACL], #4452; [NAB], 647; [EBW]; [BOW]: pl. 16, f. 2; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 309; Dunford, 2009, Insecta Mundi 90: 7; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [BAMONA]
Larva on Viola pedtifida, V. papilionacea, V. lanceolata, V. pedata, V. sagittata, V. sororia Dunford, 2009, Insecta Mundi 90: 10
- S. i. idalia New York, New Jersey, ...
  - Argynnis astarte Fisher, 1858; Proc. Acad. Nat. Sci. Philad. 10: 179 (preocc. Argynnis astarte Doubleday, 1847); TL: Nova Caesarea, Succasuma Plains, nr Schooley's Mtn, Morris Co.
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 309
    = Speyeria idalia; Dunford, 2009, Insecta Mundi 90: 7
  - Argynnis astaroth Fisher, 1859; Proc. Acad. Nat. Sci. Philad. 11: 352 (repl. Argynnis astarte Fisher, 1859); TL: Nova Caesarea, Succasuma Plains, nr Schooley's Mtn, Morris Co.
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 309
    = Speyeria idalia; Dunford, 2009, Insecta Mundi 90: 7
  - Argynnis idalia infumata Oberthür, 1912; Étud. Lépid. Comp. 6: 315, pl. 104, f. 969; TL: Newark, N.J.
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 310
    = Speyeria idalia; Dunford, 2009, Insecta Mundi 90: 7
  - Argynnis idalia tr.f. dolli Gunder, 1927; Can. Ent. 59 (12): 286; TL: Richmond Hill, Long Island, N.Y.
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 310
    = Speyeria idalia; Dunford, 2009, Insecta Mundi 90: 7
  - Argynnis idalia f. pallida Eisner, 1942; Zool. Meded. Leiden 24 (4): 123; TL: Decorah, Iowa
    
    = Speyeria idalia; Dunford, 2009, Insecta Mundi 90: 7
  - Speyeria idalia dolli; [LepIndex]; [CoL]
  - Speyeria idalia idalia; Pelham, 2008, J. Res. Lepid. 40: 309
  - Speyeria idalia infumata; [LepIndex]; [CoL]
  - Speyeria idalia pallida; [LepIndex]; [CoL]
- S. i. occidentalis Williams, 2002 Illinois, ...
  - Speyeria idalia occidentalis Williams, 2002; J. Lep. Soc. 55 (4): 146; TL: 1 mile SE of Crete, Illinois
  - Speyeria idalia occidentalis; Pelham, 2008, J. Res. Lepid. 40: 310
Speyeria nokomis (Edwards, 1862) Utah, Arizona. See [maps] Nokomis Fritillary
- Argynnis nokomis Edwards, 1862; Proc. Acad. nat. Sci. Philad. 14: 221; TL: Mt. Sneffels, Ouray Co., Colorado
- Argynnis nokomis; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 674; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13; [MPG] ; [iNaturalist] ; [BOA]
- Speyeria nokomis caerulescens; [LepIndex]; [CoL]
- Speyeria nokomis; [NACL], #4453; [NAB], 548; [EBW]; [BOW]: pl. 16, f. 8; [Opler]; [NL4A], #2289; Pelham, 2008, J. Res. Lepid. 40: 310; Dunford, 2009, Insecta Mundi 90: 15; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [BAMONA]
Larva on Viola sororia Dunford, 2009, Insecta Mundi 90: 17
- S. n. nokomis California, New Mexico, ...
  - Argynnis nitocris var. nigrocaerulea Cockerell, 1900; Ent. News 11 (10): 622; TL: Beulah, Sapello Canon, New Mexico
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 310
  - Argynnis nitocris var. nigracaerulea[sic] ab. rufescens Cockerell, 1909; Ent. Rec. J. Var. 21 (7/8): 186; TL: Beulah, Sapello Cañon
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 310
  - Argynnis nitocris nigrocaerulea; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Speyeria nokomis nigrocaerulea; [LepIndex]; [CoL]
  - Speyeria nokomis nokomis; Pelham, 2008, J. Res. Lepid. 40: 310
- S. n. nitocris (Edwards, 1874) Arizona, ...
  - Argynnis nitocris Edwards, 1874; Trans. amer. ent. Soc. 5 (1): 15; TL: White Mtns, Arizona
  - Argynnis nitocris; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Speyeria nitocris; [BOW]: pl. 16, f. 7; [LepIndex] ; [CoL]
  - Speyeria nokomis nitocris; [NACL], #4453a; Pelham, 2008, J. Res. Lepid. 40: 310
  - nitrocris[sic, nitocris]; dos Passos, 1964
- S. n. coerulescens (Holland, 1900) Mexico
  - Argynnis nitocris var. coerulescens Holland, 1900; Ent. News 11 (1): 332; TL: Mexico, Chihuahua
  - Argynnis nitocris caerulescens[sic, coerulescens]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Argynnis nitocris var. coerulescens; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 675, 3: pl. 112, f. 15-18
  - Speyeria nokomis coerulescens; [NACL], #4453b; [NL4A], #2289a; Pelham, 2008, J. Res. Lepid. 40: 310
  - caerulescens[sic, coerulescens]; McDunnough, 1938
- S. n. apacheana (Skinner, 1918) California, ...
  - Argynnis apacheana Skinner, 1918; Ent. News 29 (2): 67; TL: near Independence, Inyo Co., California
  - Argynnis apacheana ab. hermosa Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (1): 3; TL: Round Valley, Inyo Co., California
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 311
  - Argynnis apacheana; Brown, 1983, J. Lep. Soc. 37 (1): 79-80
  - Argynnis nokomis apacheana; [iNaturalist] ; [BOA]
  - Speyeria nokomis apacheana; [NACL], #4453c; Pelham, 2008, J. Res. Lepid. 40: 310; [LepIndex]; [CoL]
- S. n. wenona dos Passos & Grey, 1945 Mexico
  - Speyeria wenona dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17); TL: Cerro Potosi, Municipio de Galeana, Nuevo Leon, Mexico, 12000ft
  - Argynnis nokomis wenona; [iNaturalist] ; [BOA]
  - Speyeria nokomis wenona; [NL4A], #2289c; Pelham, 2008, J. Res. Lepid. 40: 448 (extralim.); [LepIndex]; [CoL]
- S. n. melaena Mooser & Garcia, 1980 Mexico
  - nokomis melaena Mooser & Garcia, 1980; ; TL: Mexico, Aguascalienes
  - Speyeria nokomis melaena; [NL4A], #2289b
- S. n. carsonensis Austin, 1998 Nevada
  - Speyeria nokomis carsonensis Austin, 1998; Syst. W. N. Am. Butts. (46): 573
  - Speyeria nokomis carsonensis; Pelham, 2008, J. Res. Lepid. 40: 311
- S. n. tularosa Holland, 2010 New Mexico
  - Speyeria nokomis tularosa Holland, 2010; J. Res. Lepid. 42: (79-80)
Speyeria edwardsii (Reakirt, 1866) Colorado, Utah, Nevada. See [maps] Edward's Fritillary
- Argynnis edwardsii Reakirt, 1866; Proc. Ent. Soc. Philad. 6 (1): 137; TL: Pikes Peak, Colorado
- Acidalia edwardsi montana Reuss, 1926; Dt. ent. Zs. 1926 (5): 439; TL: Montana
  - = ; Pelham, 2008, J. Res. Lepid. 40: 311; Dunford, 2009, Insecta Mundi 90: 17
- Argynnis edwardsii tr.f. edonis Gunder, 1934; Can. Ent. 66 (6): 125; TL: Beaver City, Larimor Co., Colorado
  - = ; Pelham, 2008, J. Res. Lepid. 40: 311; Dunford, 2009, Insecta Mundi 90: 17
- Argynnis edwardsii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14; [MPG] ; [iNaturalist] ; [BOA]
- Speyeria edwardsii edonis; [LepIndex]; [CoL]
- Speyeria edwardsii; [NACL], #4454; [NAB], 549; [EBW]; [BOW]: pl. 15, f. 19; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 311; Dunford, 2009, Insecta Mundi 90: 17; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [BAMONA]
Larva on Viola adunca, V. nuttallii Dunford, 2009, Insecta Mundi 90: 18

Speyeria coronis (Behr, 1864)

California. See [maps] Coronis Fritillary

Argynnis coronis Behr, 1864; Proc. ent. Soc. Philad. 3: 435; TL: California

Argynnis coronis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14; [MPG] ; [iNaturalist] ; [BOA]
Speyeria coronis; [NACL], #4455; [NAB], 550; [BOW]: pl. 15, f. 13; [Opler]; [NL4A], #2288; Pelham, 2008, J. Res. Lepid. 40: 311; Dunford, 2009, Insecta Mundi 90: 18; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [BAMONA]

identification uncertain 600x834 (~64Kb) underside USA: FR 9705 x US Route 97, (47°18'38N 120°41'48W), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

identification uncertain 600x762 (~49Kb) USA: Table Mtn, Kittitas Co., WA, 30.7.1999, Photo © Markku Savela

identification uncertain 600x619 (~54Kb) underside USA: FR9708, Table Mtn, Kittitas Co., WA, 19.7.1999, Photo © Markku Savela

Larva on Viola beckwithii, Viola douglasii, V. nuttallii, V. purpurea, V. trinervata Dunford, 2009, Insecta Mundi 90: 19

S. c. coronis
- Argynnis californica Skinner, 1917; Ent. News 28 (7): 328; TL: California, Alma, Santa Clara Co.
  - = ; Pelham, 2008, J. Res. Lepid. 40: 311
  - = Speyeria coronis; Dunford, 2009, Insecta Mundi 90: 18
- Speyeria coronis coronis; Pelham, 2008, J. Res. Lepid. 40: 311
S. c. halcyone (Edwards, 1869) Colorado
- Argynnis halcyone Edwards, 1869; Butts. N. Amer. 1: 83; TL: Colorado
- Argynnis halcyone; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria coronis halcyone; [NACL], #4455f; Pelham, 2008, J. Res. Lepid. 40: 312; [LepIndex]; [CoL]
- Speyeria halcyone; [BOW]: pl. 15, f. 20
S. c. semiramis (Edwards, 1886) WNA, California
- Argynnis semiramis Edwards, 1886; Can. Ent. 18 (4): 61; TL: California
- Argynnis semiramis tr.f. bernardensis Gunder, 1933; Can. Ent. 65 (8): 172; TL: California
  - = ; [NL4A], #2288a; Pelham, 2008, J. Res. Lepid. 40: 311
- Argynnis semiramis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
- Speyeria coronis semiramis; [NACL], #4455b; [NL4A], #2288a; Pelham, 2008, J. Res. Lepid. 40: 311; [LepIndex]; [CoL]
- Speyeria semiramis; [BOW]: pl. 16, f. 10
S. c. snyderi (Skinner, 1897) Utah
- Argynnis snyderi Skinner, 1897; Can. Ent. 29 (6): 154; TL: Salt Lake City, Utah
- Argynnis snyderi; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria coronis snyderi; [NACL], #4455e; Pelham, 2008, J. Res. Lepid. 40: 312; [LepIndex]; [CoL]
S. c. hennei (Gunder, 1934) California, ...
- Argynnis hennei Gunder, 1934; Can. Ent. 66 (6): 126; TL: nr summit, 7700ft, Mt Pinos, Frazier Mtn Park, Santa Barbara Nat. Forest, Ventura Co., California
- Speyeria coronis hennei; [NACL], #4455a; Pelham, 2008, J. Res. Lepid. 40: 311; [LepIndex]; [CoL]
S. c. carolae (dos Passos & Grey, 1942) Nevada
- Argynnis coronis carolae dos Passos & Grey, 1942; Amer. Mus. Novit. 1214: 2; TL: Charleston Park, Clark Co., Nevada
- Argynnis coronis carolae; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 21; [iNaturalist] ; [BOA]
- Speyeria carolae; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 312; Dunford, 2009, Insecta Mundi 90: 21; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [BAMONA]
- Speyeria coronis carolae; [LepIndex]; [CoL]
- Speyeria zerene carolae; [NACL], #4456e
Larva on Viola charlestonensis? Dunford, 2009, Insecta Mundi 90: 21
S. c. simaetha dos Passos & Grey, 1945 Washington, ...
- Speyeria coronis simaetha dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 2; TL: Black Canyon, Cascade Mts., nr Brewster, Washington
- Speyeria coronis simaetha; [NACL], #4455d; Pelham, 2008, J. Res. Lepid. 40: 312; [LepIndex]; [CoL]

*S. coronis* or *S. zerene*?
1600x1171 (~287Kb) upperside USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela	1600x1311 (~257Kb) underside USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela	1300x1787 (~293Kb) USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela
1500x1183 (~227Kb) USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela	1200x1598 (~310Kb) upperside USA: Idaho, Fremont Co., on Hwy 20, Sheep Falls Road (44°11'39"N 111°26'39"W), 3.8.2012, Photo © Markku Savela	900x1001 (~128Kb) USA: Idaho, Fremont Co., on Hwy 20, Sheep Falls Road (44°11'39"N 111°26'39"W), 3.8.2012, Photo © Markku Savela
1200x1455 (~228Kb) USA: Idaho, Fremont Co., on Hwy 20, Sheep Falls Road (44°11'39"N 111°26'39"W), 3.8.2012, Photo © Markku Savela

Speyeria zerene (Boisduval, 1852)

California. See [maps] Zerene Fritillary

Argynnis zerene Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 (2): 303; TL: "Yosemite"? or Agua Fria, W of Mariposa

Argynnis zerene; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14; [MPG] ; [iNaturalist] ; [BOA] ; [BOA] ; [BOA] ; [BOA]
Speyeria zerene; [NACL], #4456; [NAB], 551; [EBW]; [BOW]: pl. 16, f. 9, 11; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 313; Dunford, 2009, Insecta Mundi 90: 19; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [LEPIFORUM] ; [BAMONA]

identification uncertain 700x881 (~88Kb) underside USA: FR 9712 nr. Liberty, Lion Gulch, (47°15'47N 120°40'15W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

identification uncertain 700x904 (~73Kb) USA: FR 9705 nr. Liberty and Lion Gulch, (47°17'49N 120°39'58W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

identification uncertain 600x787 (~65Kb) underside USA: FR 9705 nr. US Route 97, (47°18'15N 120°41'24W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

Larva on Viola adunca, Viola cuneata, V. lobata, V. nuttallii, V. psychodes, V. purpurea, V. beckwithii Dunford, 2009, Insecta Mundi 90: 21

S. z. zerene E.Oregon, California
- Argynnis zerene f. conchyliatus Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 63; TL: Mt. Shasta, California
  - = ; Pelham, 2008, J. Res. Lepid. 40: 313
- Argynnis zerene ab. shastaensis Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 65; TL: Mt. Shasta, Siskiyo Co.
  - = ; Pelham, 2008, J. Res. Lepid. 40: 313
- Speyeria zerene conchyliata; [NACL], #4456a
- Speyeria zerene conchyliatus; [LepIndex]; [CoL]
- Speyeria zerene zerene; Pelham, 2008, J. Res. Lepid. 40: 313
S. z. monticola (Behr, 1863) California, ...
- Argynnis monticola Behr, 1863; Proc. Calif. Acad. Sci. 3 (1): 84; TL: Yosemite Valley
  - = Speyeria zerene; Dunford, 2009, Insecta Mundi 90: 19
- Argynnis monticola; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria zerene monticola; Pelham, 2008, J. Res. Lepid. 40: 313
S. z. behrensii (Edwards, 1869) California, ...
- Argynnis behrensii Edwards, 1869; Trans. amer. ent. Soc. 2 (4): 370; TL: Menocino, California
- Argynnis behrensii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria zerene behrensii; [NACL], #4456g; Pelham, 2008, J. Res. Lepid. 40: 312; [LepIndex]; [CoL]
S. z. bremnerii (Edwards, 1872) inland (Vancouver Island - W.Washington - W.Oregon)
- Argynnis bremnerii Edwards, 1872; Trans. amer. ent. Soc. 4: 63; TL: San Juan I.
- Argynnis bremneri v. sordida Wright, 1905; Butts. West Coast US: 133; TL: Puget Sound, Washington
  - = ; Pelham, 2008, J. Res. Lepid. 40: 312
- Argynnis bremnerii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria zerene bremnerii; [NACL], #4456i; Pelham, 2008, J. Res. Lepid. 40: 312; [LepIndex]; [CoL]
- Speyeria zerene sordida; [NACL], #4456c; [LepIndex]; [CoL]
S. z. hippolyta (Edwards, 1879) coastal (California, W.Oregon, W.Washington)
- Argynnis hippolyta Edwards, 1879; Can. Ent. 11 (5): 81; TL: Oregon; Northern California
- Argynnis hippolyta; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13; [iNaturalist]
- Speyeria hippolyta; [BOW]: pl. 15, f. 21
- Speyeria zerene hippolyta; [NACL], #4456f; McCorkle & Hammond, 1988, J. Lep. Soc. 42 (3): 184-195; Pelham, 2008, J. Res. Lepid. 40: 312; [LepIndex]; [CoL]
Larva on Viola adunca, ;
S. z. atossa (Edwards, 1890) California
- Argynnis atossa Edwards, 1890; Butts N. Amer. 3: [127], pl. 8, f. 4-5
- Argynnis atossa f. tejonica Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 68; TL: Tejon region
  - = Speyeria adiaste atossa; Pelham, 2008, J. Res. Lepid. 40: 319
- Argynnis atossa; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
- Argynnis zerene atossa; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19; [iNaturalist] ; [BOA]
- Speyeria adiaste atossa; Pelham, 2008, J. Res. Lepid. 40: 319
- Speyeria atossa; [BOW]: pl. 15, f. 8
- Speyeria egleis atossa; [NACL], #4458c; [LepIndex]; [CoL]
S. z. platina (Skinner, 1897) Utah, Idaho
- Argynnis platina Skinner, 1897; Can. Ent. 29 (6): 154; TL: Ogden, Utan; Beaver Canon, Idaho
- Argynnis pfoutsi Gunder, 1933; Can. Ent. 65 (8): 171; TL: Mt. Loafer, nr Payson, Utah
  - = ; Grey, 1989, J. Lep. Soc. 43 (1): 3; Pelham, 2008, J. Res. Lepid. 40: 314
- Argynnis platina; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria zerene pfoutsi; [NACL], #4456n; [LepIndex]; [CoL]
- Speyeria zerene platina; [NACL], #4456m; Pelham, 2008, J. Res. Lepid. 40: 314; [LepIndex]; [CoL]
S. z. malcolmi (Comstock, 1920) California
- Argynnis malcolmi Comstock, 1920; Southwest Sci. Bull. [1]: 4, pl. [1], f. 4-6; TL: Mammoth, California
- Argynnis montivaga r. malcolmi tr.f. sineargentatus Gunder, 1927; Can. Ent. 59 (12): 286; TL: Mammoth, Mono Co., California
  - = ; Pelham, 2008, J. Res. Lepid. 40: 313
- Speyeria zerene malcolmi; [NACL], #4456d; Pelham, 2008, J. Res. Lepid. 40: 313; [LepIndex]; [CoL]
S. z. picta (McDunnough, 1924) British Columbia, ...
- Argynnis halcyone r. picta McDunnough, 1924; Can. ent. 56 (2): 43; TL: Aspen Grove, B. C.
- Argynnis garretti Gunder, 1932; Can. Ent. 64 (12): 282; TL: Cranbrook, B. C.
  - = ; [NACL], #4456j; Pelham, 2008, J. Res. Lepid. 40: 314
- Speyeria zerene garretti; [NACL], #4456k; [LepIndex]; [CoL]
- Speyeria zerene picta; [NACL], #4456j; Pelham, 2008, J. Res. Lepid. 40: 314; [LepIndex]; [CoL]
S. z. gunderi (Comstock, 1925) California, ...
- Argynnis gunderi Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 67; TL: Modoc Co., California
- Speyeria zerene cynna dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 4; TL: Humboldt Nat. Forest, Ruby Valley, Elko Co., NV
  - = ; Grey, 1989, J. Lep. Soc. 43 (1): 3; Pelham, 2008, J. Res. Lepid. 40: 313
- Speyeria coronis gunderi; [NACL], #4455c; [LepIndex]; [CoL]
- Speyeria zerene cynna; [NACL], #4456o; [LepIndex]; [CoL]
- Speyeria zerene gunderi; Grey, 1989, J. Lep. Soc. 43 (1): 3; Pelham, 2008, J. Res. Lepid. 40: 313
S. z. myrtleae dos Passos & Grey, 1945 California
- Speyeria zerene myrtleae dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 3; TL: San Mateo, California
- Speyeria zerene myrtleae; [NACL], #4456h; Pelham, 2008, J. Res. Lepid. 40: 313; [LepIndex]; [CoL]
S. z. sinope dos Passos & Grey, 1945 Colorado, ...
- Speyeria zerene sinope dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 4; TL: Estes Park area, Rocky Mt. National Park, Colorado
- Speyeria zerene sinope; [NACL], #4456l; Pelham, 2008, J. Res. Lepid. 40: 314; [LepIndex]; [CoL]
S. z. gloriosa Moeck, 1957 Oregon, ...
- Speyeria zerene gloriosa Moeck, 1957; Milwaukee Ent. Soc., Special Paper: 21, 24; TL: Illinois River Valley, Josephine Co., SW.Oregon
- Speyeria zerene gloriosa; [NACL], #4456b; Pelham, 2008, J. Res. Lepid. 40: 312; [LepIndex]; [LepIndex]; [CoL]
S. z. puntareyes Emmel, Emmel & Mattoon, 1998 California
- Speyeria zerene puntareyes Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (36): 451; TL: California, Marin Coo.; Point Reyes Peninsula
- Speyeria zerene puntareyes; Pelham, 2008, J. Res. Lepid. 40: 313
S. z. sonomensis Emmel, Emmel & Mattoon, 1998 California
- Speyeria zerene sonomensis Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (36): 455
- Speyeria zerene sonomensis; Pelham, 2008, J. Res. Lepid. 40: 313
S. z. sitka Hammond, Harry & McCorkle, 2001 Alaska
- Speyeria zerene sitka Hammond, Harry & McCorkle, 2001; Butts. B. C.: 274
- Speyeria zerene sitka; Pelham, 2008, J. Res. Lepid. 40: 312

Speyeria callippe (Boisduval, 1852) -. See [maps] Callippe Fritillary
- Argynnis callippe Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 (2): 302; TL: San Francisco
- Argynnis callippe; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14; [MPG] ; [iNaturalist] ; [BOA] ; [BOA] ; [BOA]
- Speyeria callippe; [NACL], #4457; [NAB], 552; [BOW]: pl. 15, f. 11; [Opler]; [NL4A], #2287; Pelham, 2008, J. Res. Lepid. 40: 314; Dunford, 2009, Insecta Mundi 90: 22; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [BAMONA]
Larva on Viola beckwithii, Viola douglasii, V. nuttallii, V. pedunculata, V. purpurea, V. purpurea quercetorum Dunford, 2009, Insecta Mundi 90: 23
- S. c. callippe California, ...
  - Speyeria callippe callippe; Pelham, 2008, J. Res. Lepid. 40: 314
- S. c. rupestris (Behr, 1863) California, ...
  - Argynnis rupestris Behr, 1863; Proc. Calif. Acad. Sci. 3 (1): 84; TL: California
  - ?irene Edwards
    
    = Argynnis rupestris; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
  - Argynnis rupestris; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria callippe rupestris; [NACL], #4457e; Pelham, 2008, J. Res. Lepid. 40: 315; [LepIndex]; [CoL]
- S. c. juba (Boisduval, 1869) California, Nevada
  - Argynnis juba Boisduval, 1869; Ann. Soc. ent. Belg. 12: 60
  - Argynnis laura Edwards, 1879; Can. Ent. 11 (3): 49; TL: Nevada
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 315
  - Speyeria callippe sierra dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 5; TL: Gold Lake, Sierra Co., California
    
    = ?; Grey, 1989, J. Lep. Soc. 43 (1): 4
    = ; Pelham, 2008, J. Res. Lepid. 40: 315
  - Argynnis laura; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria callippe juba; [NACL], #4457f; Pelham, 2008, J. Res. Lepid. 40: 315; [LepIndex]; [CoL]
  - Speyeria callippe laura; [NACL], #4457g; [LepIndex]; [CoL]
  - Speyeria callippe sierra; [NACL], #4457h; [LepIndex]; [CoL]
- S. c. nevadensis (Edwards, 1870) Utah, Nevada, Wyoming
  - Argynnis nevadensis Edwards, 1870; Trans. amer. ent. Soc. 3 (1): 14; TL: valleys of the Sierra, nr Viriginia City
  - Argynnis nevadensis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
  - Speyeria callippe nevadensis; [NACL], #4457i; Pelham, 2008, J. Res. Lepid. 40: 316
  - Speyeria nevadensis; [BOW]: pl. 16, f. 6; [LepIndex] ; [CoL]
- S. c. inornata (Edwards, 1872) California
  - Argynnis inornata Edwards, 1872; Trans. amer. ent. Soc. 4: 64; TL: "at Downieville, California"
  - Argynnis inornata; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria callippe inornata; Pelham, 2008, J. Res. Lepid. 40: 315
- S. c. meadii (Edwards, 1872) Colorado, ...
  - Argynnis meadii Edwards, 1872; Trans. amer. ent. Soc. 4: 69; TL: Turkey Creek Junction, Colorado
  - Argynnis nevadensis r. meadii tr.f. gerhardi Gunder, 1927; Can. Ent. 59 (12): 287; TL: Colorado
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 316
  - Argynnis nevadensis meadii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
  - Speyeria callippe meadii; [NACL], #4457m; Pelham, 2008, J. Res. Lepid. 40: 316
  - Speyeria nevadensis meadii; [LepIndex]; [CoL]
- S. c. macaria (Edwards, 1877) California, Nevada
  - Argynnis macaria Edwards, 1877; Field & Forest 3 (5): 86; TL: havilah, California
  - Speyeria callippe macaria; [NACL], #4457j; Pelham, 2008, J. Res. Lepid. 40: 315; [LepIndex]; [CoL]
  - Speyeria macaria; [BOW]: pl. 16, f. 5
- S. c. liliana (H. Edwards, 1877) N.California
  - Argynnis liliana H. Edwards, 1877; Proc. Calif. Acad. Sci. 7 (12): 170; TL: nr St Helena, Napa Co.
  - Argynnis liliana var. baroni Edwards, 1881; Trans. Amer. Ent. Soc. 9: 3; TL: Northern California
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 314
  - Argynnis liliana baroni; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
  - Argynnis liliana; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
  - Speyeria callippe liliana; [NACL], #4457b; Pelham, 2008, J. Res. Lepid. 40: 314; [LepIndex]; [CoL]
- S. c. laurina (Wright, 1905) California
  - Arynnis laurina Wright, 1905; Butts. West Coast US: 53; TL: Southern California
  - Argynnis wrightii Wright, 1905; Butts. West Coast US: 139
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 315 (nom. nud.)
  - Speyeria callippe laurina; [NACL], #4457k; Pelham, 2008, J. Res. Lepid. 40: 315; [LepIndex]; [CoL]
- S. c. semivirida (McDunnough, 1924) British Columbia
  - Argynnis callippe r. semivirida McDunnough, 1924; Can. ent. 56 (2): 42; TL: Aspen Grove, B.C.
  - Argynnis callippe semivirida; [iNaturalist] ; [BOA]
  - Speyeria callippe semivirida; [NACL], #4457c; Pelham, 2008, J. Res. Lepid. 40: 316; [LepIndex]; [CoL]
- S. c. calgariana (McDunnough, 1924) Alberta
  - Argynnis nevadensis r. calgariana McDunnough, 1924; Can. ent. 56 (2): 42; TL: Calgary .. Alta
  - Speyeria calgariana; [BOW]: pl. 15, f. 10
  - Speyeria callippe calgariana; [NACL], #4457o; Pelham, 2008, J. Res. Lepid. 40: 316
  - Speyeria nevadensis calgariana; [LepIndex]; [CoL]
- S. c. comstocki (Gunder, 1925) California
  - Argynnis calippe[sic] var. comstocki Gunder, 1925; Ent. News 36: 8; TL: California
  - Argynnis comstocki tr.f. creelmani Gunder, 1934; Can. Ent. 66 (6): 127; TL: California
    
    = ; [NL4A], #2287a; Pelham, 2008, J. Res. Lepid. 40: 315
  - Speyeria callippe comstocki; [NACL], #4457a; [NL4A], #2287a; Pelham, 2008, J. Res. Lepid. 40: 315; [LepIndex]; [CoL]
- S. c. gallatini (McDunnough, 1929) Montana
  - Argynnis nevadenis var. gallatini McDunnough, 1929; Can. ent. 61 (5): 106; TL: Elkohn Ranch, Upper Gallatin Canyon, Montana
  - Speyeria callippe gallatini; [NACL], #4457n; Pelham, 2008, J. Res. Lepid. 40: 316
  - Speyeria nevadensis gallatini; [LepIndex]; [CoL]
- S. c. elaine dos Passos & Grey, 1945 Oregon
  - Speyeria callippe elaine dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 5; TL: Butte Falls, Oregon
  - Speyeria callippe elaine; [NACL], #4457d; Pelham, 2008, J. Res. Lepid. 40: 316; [LepIndex]; [CoL]
- S. c. harmonia dos Passos & Grey, 1945 Nevada
  - Speyeria callippe harmonia dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 6; TL: Mt. Wheeler, Snake Range, nr. Utah border, NV, 8000ft
  - Speyeria callippe harmonia; [NACL], #4457l; Pelham, 2008, J. Res. Lepid. 40: 316
  - Speyeria nevadensis harmonia; [LepIndex]; [CoL]
- S. c. shasta Emmel, Emmel & Mattoon, 1998 California
  - Speyeria callippe shasta Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 149; TL: California, Shasta Co.
  - Speyeria callippe shasta; Pelham, 2008, J. Res. Lepid. 40: 316
- S. c. adiasteoides Emmel, Emmel & Mattoon, 1998 California
  - Speyeria callippe adiasteoides Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 150; TL: California, Mendocino Co.. Cahto Peak summit, 4000-4200ft
  - Speyeria callippe adiasteoides; Pelham, 2008, J. Res. Lepid. 40: 314
- S. c. hagemani Emmel, Emmel & Mattoon, 1998 California
  - Speyeria callippe hagemani Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 150; TL: California, Shata Co., Shingletown, 3500'
  - Speyeria callippe hagemani; Pelham, 2008, J. Res. Lepid. 40: 316
- S. c. chilcotinensis Guppy & Shepard, 2001 British Columbia
  - Speyeria callippe chilcotinensis Guppy & Shepard, 2001; Butts. B.C.: 276; TL: BC, Riske Creek
  - Speyeria callippe chilcotensis[sic, chilcotinensis]; Pelham, 2008, J. Res. Lepid. 40: 316
Speyeria egleis (Behr, 1862) -. See [maps] Egleis Fritillary
- Argynnis egleis Behr, 1862; Proc. Cal. Acad. Sci. 2: 174 (Boisd.)
- Argynnis egleis Boisduval, 1869; Ann. Soc. ent. Belg. 12: 59
- ?nenoquis Reakirt
  - = Argynnis egleis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
- Argynnis egleis; [MPG] ; [iNaturalist] ; [BOA] ; [BOA]
- Argynnis egleis Boisduval, 1869; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
- Speyeria egleis; [NACL], #4458; [NAB], 553; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 317; Dunford, 2009, Insecta Mundi 90: 23; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [BAMONA]
Larva on Viola adunca, V. nuttallii, V. ocellata, Viola pupurea, V. purpurea integrifolia, Viola purpurea venosa, V. walteri, Festuca ovina, Potentilla Dunford, 2009, Insecta Mundi 90: 24
- S. e. egleis California
  - Argynnis montivaga Behr, 1863; Proc. Calif. Acad. Sci. 3 (1): 84; TL: California
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 317
    = Speyeria egleis; Dunford, 2009, Insecta Mundi 90: 23
  - Argynnis egleis Boisduval, 1869; Ann. Soc. ent. Belg. 12: 59
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 317
  - Argynnis astarte Edwards, 1864; Proc. ent. Soc. Philad. 3: 434 (preocc. Arynnis astarte Doubleday, 1847)
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 317
    = Speyeria egleis; Dunford, 2009, Insecta Mundi 90: 23
  - Argynnis montivaga ab. mammothi Gunder, 1924; Ent. News 35 (5): 157, pl. 2, f. H, D
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 317
    = Speyeria egleis; Dunford, 2009, Insecta Mundi 90: 23
  - Argynnis montivaga tr.f. boharti Gunder, 1929; Bull. Brooklyn ent. Soc. 24 (5): 326, pl. 31, f. 5
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 317
    = Speyeria egleis; Dunford, 2009, Insecta Mundi 90: 23
  - Argynnis egleis; [BOA]
  - Argynnis montivaga; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria egleis egleis; Pelham, 2008, J. Res. Lepid. 40: 317
- S. e. oweni (Edwards, 1892) S.Oregon - California
  - Argynnis oweni Edwards, 1892; Can. Ent. 24 (5): 105; TL: Mt. Shasta, Sikiyou Co., California
  - Argynnys hippolyta oweni; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Speyeria egleis oweni; [NACL], #4458e; Hammond & Dornfeld, 1983, J. Lep. Soc. 37 (2): (note) , f. 3; Pelham, 2008, J. Res. Lepid. 40: 317; [LepIndex]; [CoL]
- S. e. utahensis (Skinner, 1919) Utah
  - Argynnis utahensis Skinner, 1919; Ent. News 30 (8): 216; TL: City Creek Canyon, Salt Lake City, Utah
  - Speyeria egleis utahensis; [NACL], #4458i; Pelham, 2008, J. Res. Lepid. 40: 318; [LepIndex]; [CoL]
- S. e. tehachapina (Comstock, 1920) California
  - Argynnis montivaga var. tehachapina Comstock, 1920; Southwest Sci. Bull. [1]: 6; TL: highest peak in Tehachapi Mts, California
  - Speyeria egleis tehachapina; [NACL], #4458d; Pelham, 2008, J. Res. Lepid. 40: 317; [LepIndex]; [CoL]
- S. e. macdunnoughi (Gunder, 1932) Montana
  - Argynnis macdunnoughi Gunder, 1932; Can. Ent. 64 (12): 280; TL: Elkhorn Ranch Resort, Gallatin Co., Montana
  - Speyeria egleis macdunnoughi; [NACL], #4458g; Pelham, 2008, J. Res. Lepid. 40: 318; [LepIndex]; [CoL]
- S. e. albrighti (Gunder, 1932) Montana
  - Argynnis albrighti Gunder, 1932; Can. Ent. 64 (12): 281; TL: Highwood Mts., Choteau Co., Montana
  - Speyeria egleis albrighti; [NACL], #4458h; Pelham, 2008, J. Res. Lepid. 40: 318; [LepIndex]; [CoL]
- S. e. linda (dos Passos & Grey, 1942) Idaho
  - Argynnis utahensis linda dos Passos & Grey, 1942; Amer. Mus. Novit. 1214: 1; TL: Heyburn Peak, Sawtooth-Boise, ID, 9500-10000ft
  - Speyeria egleis linda; [NACL], #4458f; Pelham, 2008, J. Res. Lepid. 40: 318; [LepIndex]; [CoL]
- S. e. secreta dos Passos & Grey, 1945 Colorado
  - Speyeria montiviga[sic] secreta dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 7; TL: Estes Park area, Rocky Mt. National Park, Colorado, 8000ft
  - Speyeria egleis secreta; [NACL], #4458k; Pelham, 2008, J. Res. Lepid. 40: 318; [LepIndex]; [CoL]
- S. e. toiyabe Howe, 1975 Nevada
  - Speyeria egleis toiyabe Howe, 1975; Butterflies of North America: 237; TL: Kingston Canyon, Toiyabe Range, Lander Co., Nevada
  - Speyeria egleis toiyabe; [NACL], #4458j; Pelham, 2008, J. Res. Lepid. 40: 318; [LepIndex]; [CoL]
- S. e. moecki Hammond & Dornfeld, 1983 Oregon
  - Speyeria egleis moecki Hammond & Dornfeld, 1983; J. Lep. Soc. 37 (2): 115, f. 2; TL: Skookum Mdw., Walker Rim, Klamath Co., Oregon
  - Speyeria egleis moecki; Pelham, 2008, J. Res. Lepid. 40: 318
  Larva on Viola purpurea Hammond & Dornfeld, 1983, J. Lep. Soc. 37 (2): 120
- S. e. mattooni Emmel & Emmel, 1998 California
  - Speyeria egleis mattooni Emmel & Emmel, 1998; Syst. W. N. Am. Butts. (33): 438; TL: California, Siskiyou Co., Dry Lake Mtn, US FSR 40S01
  - Speyeria egleis mattooni; Pelham, 2008, J. Res. Lepid. 40: 317
- S. e. yolaboli Emmel & Emmel, 1998 California
  - Speyeria egleis yolaboli Emmel & Emmel, 1998; Syst. W. N. Am. Butts. (33): 439; TL: California, Tehama Co., South Yolla Bolly Mtn
  - Speyeria egleis yolaboli; Pelham, 2008, J. Res. Lepid. 40: 317
- S. e. reidi Austin, 1998 Nevada
  - Speyeria egleis reidi Austin, 1998; Syst. W. N. Am. Butts. (46): 574; TL: Nevada, White Pine Co., Snake Range, Wheeler Peak Campground, 3033m
  - Speyeria egleis reidi; Pelham, 2008, J. Res. Lepid. 40: 318
Speyeria adiaste (Edwards, 1864) California. See [maps] Unsilverered Fritillary Adiaste Fritillary
- Argynnis adiaste Edwards, 1864; Proc. ent. Soc. Philad. 3: 436; TL: California
- Argynnis adiaste; [MPG] ; [iNaturalist] ; [BOA]
- Argynnis adraste[sic, adiaste]; Kirby, 1871, 160
- Speyeria adiaste; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 318; Dunford, 2009, Insecta Mundi 90: 24; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [BAMONA]
- ?Argynnis adiaste Behr, 1862; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
Larva on Viola purpurea quercetorum, V. ocellata? Dunford, 2009, Insecta Mundi 90: 25
- S. a. adiaste
  - Argynnis adiante Boisduval, 1869; Ann. Soc. ent. Belg. 12: 61; TL: California
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 318
  - Argynnis adianthe; Barnes & McDunnough, 1917, 8 (missp.)
  - Speyeria adiante; [BOW]: pl. 15, f. 4
  - Speyeria adiaste adiaste; Pelham, 2008, J. Res. Lepid. 40: 318
  - Speyeria egleis adiaste; [NACL], #4458a
- S. a. clemencei (Comstock, 1925) California
  - Argynnis adiaste r. clemencei Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 67; TL: Atascadero, California
  - Argynnis adiaste clemencei; [iNaturalist] ; [BOA]
  - Speyeria adiaste clemencei; Pelham, 2008, J. Res. Lepid. 40: 319
  - Speyeria egleis clemencei; [NACL], #4458b; [LepIndex]; [CoL]

Speyeria atlantis (Edwards, 1862)

ENA. See [maps] Atlantis Fritillary

Argynnis atlantis Edwards, 1862; Proc. Acad. nat. Sci. Philad. 14: 54

Argynnis atlantis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13; [MPG] ; [iNaturalist] ; [BOA]
Speyeria atlantis; [NACL], #4459; [NAB], 554; [BOW]: pl. 15, f. 7; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 25; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [LEPIFORUM] ; [BAMONA]

identification uncertain 700x795 (~54Kb) underside USA: FR 9705 nr. US Route 97, (47°18'15N 120°41'24W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

identification uncertain 700x976 (~102Kb) underside USA: FR 9705 x US Route 97, (47°18'38N 120°41'48W), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

Larva on Viola septentrionalis, Viola sororia affinis, V. adunca, V. candensis Dunford, 2009, Insecta Mundi 90: 27

S. a. atlantis Maine, ...
- Argynnis atlantis ab. chemo Scudder, 1889; Butts eastern U.S. Canada 1: 573; TL: Lake Chemo
  - = ; Pelham, 2008, J. Res. Lepid. 40: 319
  - = Speyeria atlantis; Dunford, 2009, Insecta Mundi 90: 25
- Speyeria atlantis atlantis; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 27
S. a. canadensis (dos Passos, 1935) Newfoundland, ...
- Argynnis atlantis canadensis dos Passos, 1935; Can. Ent. 67 (4): 85; TL: Doyles Station, Newfoundland
  - = Speyeria atlantis; Dunford, 2009, Insecta Mundi 90: 25
- Argynnis atlantis canadensis; [iNaturalist] ; [BOA]
- Speyeria atlantis canadensis; [NACL], #4459a; Pelham, 2008, J. Res. Lepid. 40: 319; [LepIndex]; [CoL]
S. a. hollandi (Chermock & Chermock, 1940) Manitoba, British Columbia
- Argynnis atlantis hollandi Chermock & Chermock, 1940; Can. Ent. 72 (4): 82; TL: Riding Mtns, Manitoba
- Argynnis atlantis hollandi; [iNaturalist] ; [BOA]
- Speyeria atlantis hollandi; [NACL], #4459b; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 28; [LepIndex]; [CoL]
S. a. sorocko Scott, Kondla & Spomer, 1998 Colorado
- Speyeria atlantis sorocko Scott, Kondla & Spomer, 1998; Papilio (N.S) 8: 13; TL: several km NE of Mt. Jude, 9100ft, Clear Creek Co., Colorado
- Speyeria atlantis sorocko; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 28
Larva on Viola sororia affinis, V. scopulorum, V. canadensis Dunford, 2009, Insecta Mundi 90: 29
S. a. pahasapa Spomer, Scott & Kondla, 1998 South Dakota
- Speyeria atlantis pahasapa Spomer, Scott & Kondla, 1998; Papilio (N.S) 8: 13; TL: Deerfield Reservoir, Pennington Co., South Dakota
- Speyeria atlantis pahasapa; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 29

Speyeria hesperis (Edwards, 1864) Colorado, Utah, Montana. See [maps]
- Argynnis hesperis Edwards, 1864; Proc. ent. Soc. Philad. 2 (4): 502; TL: Rocky Mountains
- Argynnis hesperis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14; [MPG] ; [iNaturalist] ; [BOA]
- Speyeria hesperis; [BOW]: pl. 16, f. 1; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 29; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [BAMONA]
Larva on Viola canadensis var. scopulorum, V. adunca, V. sororia affinis, V. rydbergii, V. adunca bellidifolia, V. nuttallii, V. purpurea Dunford, 2009, Insecta Mundi 90: 30
- S. h. hesperis Colorado
  - Speyeria atlantis hesperis; [NACL], #4459c; [LepIndex]; [CoL]
  - Speyeria hesperis hesperis; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 29
- S. h. beani (Barnes & Benjamin, 1926) Alberta
  - Dryas atlantis r. beani Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 92; TL: Banff, Alberta
  - Speyeria atlantis beani; [NACL], #4459r; [LepIndex]; [CoL]
  - Speyeria hesperis beani; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 31
- S. h. lurana dos Passos & Grey, 1945 South Dakota
  - Speyeria atlantis lurana dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 8; TL: Harney Peak, Black Hills, SD
  - Speyeria atlantis lurana; [NACL], #4459p; [LepIndex]; [CoL]
  - Speyeria hesperis lurana; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 34
- S. h. hutchinsi dos Passos & Grey, 1947 Montana
  - Speyeria atlantis hutchinsi dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 20; TL: Twp 2N Range 1W, Jefferson Co., Montana
    
    = Speyeria hesperis beani; Dunford, 2009, Insecta Mundi 90: 31
  - Speyeria atlantis beani tr.f. hutchinsi Gunder, 1932; Can. Ent. 64 (12): 280; TL: Twp 2N Range 1W, Jefferson Co., Montana
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 320
  - Speyeria atlantis hutchinsi; [NACL], #4459q; [LepIndex]; [CoL]
  - Speyeria hesperis hutchinsi; Pelham, 2008, J. Res. Lepid. 40: 320
- S. h. helena dos Passos & Grey, 1957 Alberta, British Columbia, Manitoba
  - Argynnis lais Edwards, 1883; Can. Ent. 15 (11): 209 (preocc. Argynnis lais Scudder, 1875); TL: "Red Deer River"
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 31
  - Argynnis lais tr.f. dennisi Gunder, 1927; Can. Ent. 59 (12): 287; TL: Beulah, Manitoba
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 31
  - Speyeria atlantis dennisi dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 20; TL: Beulah, Manitoba
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 31
  - Speyeria (Speyeria) atlantis helena dos Passos & Grey, 1957; J. N.Y. Ent. Soc. 63: 95 (repl. Argynnis lais Edwards, 1883)
  - Argynnis lais; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Speyeria atlantis dennisi; [NACL], #4459t
  - Speyeria atlantis helena; [LepIndex]; [CoL]
  - Speyeria atlantis lais; [NACL], #4459s
  - Speyeria electa dennisi; [LepIndex]; [CoL]
  - Speyeria hesperis helena; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 31
  - Speyeria lais; [BOW]: pl. 16, f. 3
- S. h. ratonensis Scott, 1981 New Mexico
  - Speyeria atlantis ratonensis Scott, 1981; Papilio (n.s.) 1: 1; TL: Raton Mesa, Colfax Co., New Mexico
  - Speyeria hesperis ratonensis; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 33
  Larva on Viola canadensis var. scopulorum Dunford, 2009, Insecta Mundi 90: 33
- S. h. brico Kondla, Scott & Spomer, 1998 British Columbia
  - Speyeria hesperis brico Kondla, Scott & Spomer, 1998; Papilio (N.S) 8: 14; TL: Castle Creek FS Rd Km 23.5, Cariboo Mts, British Columbia
  - Speyeria hesperis brico; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 32
Speyeria irene (Boisduval, 1869) California, .... See [maps]
- Argynnis egleis var. irene Boisduval, 1869; Ann. Soc. ent. Belg. 12: 60
- Argynnis (Speyeria) irene; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 17
- Argynnis irene; [MPG] ; [BOA]
- Argynnis rupestris irene; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
- Speyeria atlantis irene; [NACL], #4459n; [LepIndex]; [CoL]
- Speyeria hesperis irene; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 34
Larva on Viola purpurea Dunford, 2009, Insecta Mundi 90: 35
- S. i. dodgei (Gunder, 1931) Oregon
  - Argynnis dodgei Gunder, 1931; Bull. South. Calif. Acad. Sci. 30 (2): 46; TL: Diamond Lake, Dougla Co., Oregon
  - Argynnis irene dodgei; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19; [BOA]
  - Speyeria atlantis dodgei; [NACL], #4459m; [LepIndex]; [CoL]
  - Speyeria hesperis dodgei; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 37
  Larva on Violla bellidifolia Dunford, 2009, Insecta Mundi 90: 38
- S. i. cottlei (Comstock, 1925) California
  - Argynnis cottlei Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 64; TL: nr Alturas, Modoc Co., California
  - Argynnis irene cottlei; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19; [BOA]
  - Speyeria hesperis cottlei; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 35
- S. i. hanseni (Emmel, Emmel & Mattoon, 1998) California
  - Speyeria atlantis hanseni Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 152; TL: California, Tehama Co., Anthony Peak on Covelo Rd
  - Argynnis irene hanseni; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19; [BOA]
  - Speyeria hesperis hanseni; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 36
Speyeria nausicaa (Edwards, 1874) Arizona, .... See [maps]
- Argynnis nausicaa Edwards, 1874; Trans. amer. ent. Soc. 5 (2): 104; TL: "Rocky Canon, Arizona"
- Argynnis aphrodite f. arizonensis Elwes, 1889; Trans. ent. Soc. Lond. 1889: 546; TL: Arizona
  - = Speyeria hesperis nausicaa; Pelham, 2008, J. Res. Lepid. 40: 322; Dunford, 2009, Insecta Mundi 90: 42
- Argynnis (Speyeria) nausicaa; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 17
- Argynnis nausicaa; [MPG] ; [BOA]
- Speyeria atlantis nausicaa; [NACL], #4459f; [LepIndex]; [CoL]
- Speyeria hesperis nausicaa; Pelham, 2008, J. Res. Lepid. 40: 322; Dunford, 2009, Insecta Mundi 90: 42
- S. n. elko Austin, 1983 Nevada (Elko Co.)
  - Speyeria atlantis elko Austin, 1983; J. Lep. Soc. 37 (3): 244, f. 1-2; TL: Nevada, Elko Co., Owybee River Valley, Wild Horse Creek Camping, ca. 10mi S of Mountain City
  - Argynnis nausicaa elko; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19; [BOA]
  - Speyeria hesperis elko; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 38
- S. n. greyi Moeck, 1950 Nevada
  - Speyeria atlantis greyi Moeck, 1950; Ent. News 61 (3): 61; TL: Lamoille Canyon, Ruby Mtns, Elko Co., Nevada
  - Argynnis nausicaa greyi; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19; [BOA]
  - Speyeria atlantis greyi; [NACL], #4459j; [LepIndex]; [CoL]
  - Speyeria hesperis greyi; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 33
- S. n. viola dos Passos & Grey, 1945 Idaho
  - Speyeria atlantis viola dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 10; TL: Trail Creek, Sawtooth Mts., ID, 7400ft
  - Argynnis nausicaa viola; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19; [BOA]
  - Speyeria atlantis viola; [NACL], #4459l; [LepIndex]; [CoL]
  - Speyeria hesperis viola; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 37
- S. n. tetonia (dos Passos & Grey, 1945) Wyoming, Utah
  - Speyeria atlantis wasatchia dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 8; TL: Payson Canyon, Payson, Utah
    
    = Speyeria hesperis tetonia; Pelham, 2008, J. Res. Lepid. 40: 320
  - Speyeria hydaspe conquista dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17); TL: Little Tesuque Canyon, nr Sante Fe, New Mexico, 8000ft
    
    = Speyeria hydaspe rhodope; Pelham, 2008, J. Res. Lepid. 40: 323
    = Argynnis nausicaa tetonia; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19
  - Speyeria atlantis tetonia dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 9; TL: Teton Mts., WY
  - Argynnis nausicaa tetonia; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19; [BOA]
  - Speyeria atlantis tetonia; [NACL], #4459k; [LepIndex]; [CoL]
  - Speyeria atlantis wasatchia; [NACL], #4459i; [LepIndex]; [CoL]
  - Speyeria hesperis tetonia; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 39
  - Speyeria hesperis wasatchia; Dunford, 2009, Insecta Mundi 90: 39
  - Speyeria hydaspe conquista; [NACL], #4460f; [LepIndex]; [CoL]
- S. n. chitone (Edwards, 1879) Utah, Arizona
  - Argynnis chitone Edwards, 1879; Can. Ent. 11 (5): 82; TL: Southern Utah and Arizona
  - Argynnis chitone; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
  - Argynnis nausicaa chitone; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19; [BOA]
  - Speyeria atlantis chitone; [NACL], #4459h; [LepIndex]; [CoL]
  - Speyeria hesperis chitone; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 40
- S. n. schellbachi Garth, 1949 Arizona, ...
  - Speyeria atlantis schellbachi Garth, 1949; Bull. South. Calif. Acad. Sci. 48 (1). 1-2: 1; TL: Neal Springs, North Rim, Grand Canyon National Park, Coconino Co., Arizona
  - Argynnis nausicaa schellbachi; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19; [BOA]
  - Speyeria atlantis schellbachi; [NACL], #4459g; [LepIndex]; [CoL]
  - Speyeria hesperis schellbachi; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 41
  - shellbachi; dos Passos, 1964, (missp.)
- S. n. electa (Edwards, 1878) Colorado, Montana, New Mexico
  - Argynnis electa Edwards, 1878; Field & Forest 3: 143
  - Argynnis cornelia Edwards, 1892; Can. Ent. 24 (5): 106
    
    = Speyeria hesperis electa; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 40
  - Argynnis nikias Ehrmann, 1917; Lepidopterist 1 (7): 55; TL: Temez[sic] Springs, New Mexico
    
    = Speyeria hesperis electa; Pelham, 2008, J. Res. Lepid. 40: 322; Dunford, 2009, Insecta Mundi 90: 41
  - Argynnis cornelia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Argynnis electa; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Argynnis nausicaa electa; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19; [BOA]
  - Speyeria atlantis electa; [NACL], #4459o
  - Speyeria atlantis nikias; [NACL], #4459d; [LepIndex]; [CoL]
  - Speyeria electa; [LepIndex] ; [CoL]
  - Speyeria hesperis electa; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 40
  Larva on Viola sororia affinis Dunford, 2009, Insecta Mundi 90: 41
- S. n. dorothea Moeck, 1947 New Mexico
  - Speyeria atlantis dorothea Moeck, 1947; Ent. News 58 (3): 73; TL: Sandia Peak, Sandia Mtns, New Mexico
  - Argynnis nausicaa dorothea; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19; [BOA]
  - Speyeria atlantis dorothea; [NACL], #4459e; [LepIndex]; [CoL]
  - Speyeria hesperis dorothea; Pelham, 2008, J. Res. Lepid. 40: 322; Dunford, 2009, Insecta Mundi 90: 43
  Larva on Viola sororia affinis Dunford, 2009, Insecta Mundi 90: 43
- S. n. capitatensis Holland, 1988 New Mexico
  - Speyeria atlantis capitanensis Holland, 1988; Bull. Allyn Mus. 115: 2; TL: Padilla Point, crest of Capitan Ridge, Capitan Mts, Lincoln Co., NM
  - Argynnis nausicaa capitatensis; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 19
  - Speyeria hesperis capitatensis; Pelham, 2008, J. Res. Lepid. 40: 322; Dunford, 2009, Insecta Mundi 90: 43

Speyeria hydaspe (Boisduval, 1869)

W.USA. See [maps] Hydaspe Fritillary

Argynnis hydaspe Boisduval, 1869; Ann. Soc. ent. Belg. 12: 60; TL: California

Argynnis hydaspe; [MPG] ; [iNaturalist] ; [BOA] ; [BOA]
Speyeria hydaspe; [NACL], #4460; [NAB], 555; [EBW]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 322; Dunford, 2009, Insecta Mundi 90: 44; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [BAMONA]

800x828 (~114Kb) underside USA: Taneum Creek, Wenatchee National Forest, (47°06'24N 120°54'44W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela	identification uncertain 800x758 (~67Kb) upperside USA: FR9705, Table Mtn, Kittitas Co., WA, 30.7.1999, Photo © Markku Savela	500x673 (~59Kb) ♂ USA: End of FR 6520 /Johnson Ridge, Mt. Baker Snoqualmie Nat. Forest (47°47'48N 121°15'45W), Snohomish Co., WA, 7.8.2001, Photo © Markku Savela
500x582 (~46Kb) underside USA: End of FR 6520 /Johnson Ridge, Mt. Baker Snoqualmie Nat. Forest (47°47'48N 121°15'45W), Snohomish Co., WA, 7.8.2001, Photo © Markku Savela

Larva on Viola adunca, V. glabella, V. nuttallii, V. orbiculata, V. purpurea, V. sheltonii Dunford, 2009, Insecta Mundi 90: 45

S. h. hydaspe California
- Argynnis monticola var. purpurascens H. Edwards, 1877; Proc. Calif. Acad. Sci. 7 (12): 170; TL: Siskiyou Co.
  - = ; Pelham, 2008, J. Res. Lepid. 40: 322
- Argynnis hydaspe ab. caliginosa Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 66; TL: Modoc Co., California
  - = ; Pelham, 2008, J. Res. Lepid. 40: 322
- Argynnis monticola purpurascens; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria hydaspe hydaspe; Pelham, 2008, J. Res. Lepid. 40: 322
- Speyeria hydaspe purpurascens; [NACL], #4460b; [LepIndex]; [CoL]
S. h. rhodope (Edwards, 1874) Vancouver, British Columbia
- Argynnis rhodope Edwards, 1874; Trans. amer. ent. Soc. 5 (1): 13; TL: Cariboo District, British Columbia
- Argynnis sakuntala Skinner, 1911; Ent. News 22 (3): 108; TL: Kaslo, B. C.
  - = ; Pelham, 2008, J. Res. Lepid. 40: 323
- Argynnis skinneri Holland, 1931; The Butterfly Book (1931 edn): 95, pl. 56, f. 4 (repl. Argynnis sakuntala Skinner, 1911); TL: Kaslo, B. C.
  - = ; Pelham, 2008, J. Res. Lepid. 40: 323
- Argynnis rhodope; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria hydaspe rhodope; [NACL], #4460d; Pelham, 2008, J. Res. Lepid. 40: 323; [LepIndex]; [CoL]
- Speyeria hydaspe sakuntala; [NACL], #4460e; [LepIndex]; [CoL]
S. h. viridicornis (Comstock, 1925) California
- Argynnis hydaspe f. viridicornis Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 63; TL: Greenhorn Mtns, California
- Argynnis hydaspe viridicornis; [iNaturalist] ; [BOA]
- Speyeria hydaspe viridicornis; [NACL], #4460a; Pelham, 2008, J. Res. Lepid. 40: 322; [LepIndex]; [CoL]
S. h. minor dos Passos & Grey, 1947 British Columbia
- Speyeria (Speyeria) hydaspe minor dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 21; TL: Mt. McLean, B. C.
- Argynnis rhodope tr.f. gregsoni Gunder, 1932; Can. Ent. 64 (12): 281; TL: Mt Washington, Forbidden Plateau, V.I., B.C
  - = ; Pelham, 2008, J. Res. Lepid. 40: 323
- Argynnis hydaspe rhodope f. alt. minor McDunnough, 1927; Can. ent. 59 (7): 155; TL: Mt. McLean, B. C.
  - = ; Pelham, 2008, J. Res. Lepid. 40: 323
- Speyeria hydaspe minor; [NACL], #4460c; Pelham, 2008, J. Res. Lepid. 40: 323; [LepIndex]; [CoL]
S. h. tehama Emmel, Emmel & Mattoon, 1998 California
- Speyeria hydaspe tehama Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 146; TL: California, Tehama Co., Covelo Rd, at Govenrment Flat, 5900'
- Speyeria hydaspe tehama; Pelham, 2008, J. Res. Lepid. 40: 322
S. h. davisi Emmel, Emmel & Mattoon, 1998 California
- Speyeria hydaspe davisi Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 147; TL: California, Humboldt Co., McClellan Mtn peat bog
- Speyeria hydaspe davisi; Pelham, 2008, J. Res. Lepid. 40: 323

Speyeria mormonia (Boisduval, 1869) California, Nevada. See [maps] Mormon Fritillary
- Argynnis mormonia Boisduval, 1869; Ann. Soc. ent. Belg. 12: 58
- Argynnis mormonia; [MPG] ; [iNaturalist] ; [BOA]
- Speyeria mormonia; [NACL], #4461; [NAB], 557; [EBW]; [BOW]: pl. 16, f. 4; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 323; Dunford, 2009, Insecta Mundi 90: 45; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [LepIndex] ; [CoL] ; [LEPIFORUM] ; [BAMONA]
1200x885 (~136Kb) upperside USA: Washington, Skagit Co., Sauk Mountain Trail Head & Trail, 15.8.2012, Photo © Markku Savela

Larva on Viola nuttallii, V. palustris, V. adunca, Viola adunca var. bellidifolia, V. sororia Dunford, 2009, Insecta Mundi 90: 46
- S. m. mormonia Oregon?, California
  - Argynnis astarte Edwards, 1862; Proc. Ent. Soc. Philad. 1 (7): 221 (preocc. by Doubleday, 1847)
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 325
  - Argynnis montivaga Behr, 1864; Proc. Ent. Soc. Philad. 3 (3): 435 (preocc. Argynnis montivaga Behr, 1863)
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 325
  - Argynnis arge Strecker, 1878; Synon. Cat. Macroplep.: 114
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 325
  - Argynnis eurynome arge; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria mormonia arge; [NACL], #4461f; [LepIndex]; [CoL]
  - Speyeria mormonia mormonia; Pelham, 2008, J. Res. Lepid. 40: 325
- S. m. bischoffii (Edwards, 1870) Alaska
  - Argynnis bischoffii Edwards, 1870; Trans. amer. ent. Soc. 3: 189; TL: Anchorage, Alaska
  - Argynnis bischoffii; [MPG] ; [BOA] ; [BOA]
  - Argynnis eurynome bischoffi; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria mormonia bischoffi; [LepIndex]; [CoL]
  - Speyeria mormonia bischoffii; [NACL], #4461a; Grey, 1989, J. Lep. Soc. 43 (1): 6; Pelham, 2008, J. Res. Lepid. 40: 323
- S. m. eurynome (Edwards, 1872) Wyoming, Montana, Alberta, Pacific Coast, ...
  - Argynnis eurynome Edwards, 1872; Trans. amer. ent. Soc. 4: 66
  - Argynnis clio Edwards, 1874; Trans. amer. ent. Soc. 5 (2): 106
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 324
  - Argynnis eurynome ab. eris Igel, 1922; (preocc. by Meigen, 1828)
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 324
  - Argynnis mormonia tr.f. benjamini Gunder, 1927; Can. Ent. 59 (12): 287; TL: Arangie, Idaho
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 324
  - Argynnis eurynome tr.f. brucei Gunder, 1927; Can. Ent. 59 (12): 287
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 325
  - Argynnis eurynome r. clio f. gunderi Field, 1936; J. Ent. Zool. 28 (2): 23 (preocc. by Comstock, 1925); TL: Broadwater Co., Montana
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 325
  - Speyeria mormonia eurynome ab. igeli dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 23 (repl. Argnynnis eurynome ab. eris Igel, 1922)
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 325
  - Speyeria mormonia eurynome ab. fieldi dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 23 (repl. Argynnis eurynome clio trans.f. gunderi Field, 1936)
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 325
  - Argynnis eurynome clio; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Argynnis eurynome; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria eurynome; [BOW]: pl. 15, f. 18
  - Speyeria mormonia benjamini; [LepIndex]; [CoL]
  - Speyeria mormonia eurynome; [NACL], #4461h; Pelham, 2008, J. Res. Lepid. 40: 324; [LepIndex]; [CoL]
  - eurynomes; Edwards, 1873, (missp.)
- S. m. opis (Edwards, 1874) British Columbia
  - Argynnis opis Edwards, 1874; ; TL: Bald Mt., Cariboo Dist., British Columbia
  - Argynnis bischoffi opis f. jesmondensis McDunnough, 1940; Can. ent. 72 (2): 25; TL: Jesmond, B. C.
    
    = ; Grey, 1989, J. Lep. Soc. 43 (1): 6; Pelham, 2008, J. Res. Lepid. 40: 324
  - Speyeria mormonia jesmondensis dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 22; TL: Jesmond, B. C.
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 324
  - Speyeria mormonia jesmondensis; [NACL], #4461c; [LepIndex]; [CoL]
  - Speyeria mormonia opis; [NACL], #4461b; Pelham, 2008, J. Res. Lepid. 40: 324; [LepIndex]; [CoL]
- S. m. artonis (Edwards, 1881) Nevada, ...
  - Argynnis artonis Edwards, 1881; Trans. Amer. Ent. Soc. 9: 1
  - Argynnis eurynome artonis f. rubyensis Gunder, 1932; Can. Ent. 64 (12): 283; TL: Franklin Slough, nr Wright's Ranch, Ruby Valley, Elko Co., Nevada
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 325
  - Argynnis eurynome artonis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria mormonia artonis; [NACL], #4461g; Pelham, 2008, J. Res. Lepid. 40: 325; [LepIndex]; [CoL]
- S. m. erinna (Edwards, 1883) Washington, Oregon
  - Argynnis eurynome var. erinna Edwards, 1883; Can. Ent. 15 (2): 33; TL: Spokane Falls
  - Argynnis erinna ab. cunninghami Owen, 1893; Ent. News 4 (7): 246; TL: Klamath Co., Oregon
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 324
  - Argynnis eurynome cunninghami; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Argynnis eurynome erinna; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria mormonia erinna; [NACL], #4461e; Hammond & Dornfeld, 1983, J. Lep. Soc. 37 (2): 120 (note); Pelham, 2008, J. Res. Lepid. 40: 324; [LepIndex]; [CoL]
  Larva on Viola palustris, Viola adunca var. bellidifolia Hammond & Dornfeld, 1983, J. Lep. Soc. 37 (2): 120
- S. m. washingtonia (Barnes & McDunnough, 1913) Washington
  - Argynnis bischoffi[sic] washingtonia Barnes & McDunnough, 1913; Contr. nat. Hist. Lep. N. Am. 2 (3): 95, pl. 1, f. 5; TL: Mt Ra[i]nier, 7000ft, [Pierce Co.], Wash[ington]
  - Argynnis eurynome washingtonia f. ranierensis Gunder, 1932; Can. Ent. 64 (12): 282; TL: Paradise Valley, Mount Ra[i]nier, Pierce Co., Washington
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 324
  - Speyeria mormonia washingtonia; [NACL], #4461d; Pelham, 2008, J. Res. Lepid. 40: 324; [LepIndex]; [CoL]
  - Speyeria rainierensis[sic]; [NACL], #4461
  - ranierensis; Guppy & Kondla, 2002, J. Lep. Soc. 56 (3): 181
- S. m. luski (Barnes & McDunnough, 1913) Arizona
  - Argynnis eurynome luski Barnes & McDunnough, 1913; Contr. nat. Hist. Lep. N. Am. 2 (3): 95; TL: White Mts, Arizona
  - Speyeria mormonia luski; [NACL], #4461i; Pelham, 2008, J. Res. Lepid. 40: 326; [LepIndex]; [CoL]
- S. m. obsidiana Emmel, Emmel & Mattoon, 1998 California
  - Speyeria mormonia obsidiana Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 145; TL: California, Mono Co., Sawmill Meadow, 9200', Glass Mtn
  - Speyeria mormonia obsidiana; Pelham, 2008, J. Res. Lepid. 40: 325

Unmatched external taxa

Speyeria aglaia appennicola; [LepIndex]
Speyeria aglaia; [LepIndex]
Speyeria aglaia semilocuples; [LepIndex]

27.5.2023 (20)

References:

[BAMONA] Opler, Pavulaan, Stanford & Pogue
Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA] electronic
BIOLOGIA CENTRALI-AMERICANA
[BOA] Interactive Listing of American Butterflies
Listado Interactivo de la Mariposas Americanas; Butterflies of America
[CiNii]
http://ci.nii.ac.jp/
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[EVERSMANNIA]
Eversmannia contents
[FLMNH] Florida Museum of Natural History
Bulletin of the Allyn Museum
[Heiner Ziegler]
Switzerland / Schweiz; www.pieris.ch
[JLS]
Journal of the Lepidopterists' Society; the original J. Lep. Soc. articles
[LAMAS-2005] Gerardo Lamas
Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[LepIndex] Global Lepidoptera Index
Beccaloni, G., Scoble, M., Kitching, I., Simonsen, T., Robinson, G., Pitkin, B., Hine, A., Lyal, C., Ollerenshaw, J., Wing, P., & Hobern, D. (2024). Global Lepidoptera Index. In O. Bánki, Y. Roskov, M. Döring, G. Ower, D. R. Hernández Robles, C. A. Plata Corredor, T. Stjernegaard Jeppesen, A. Örn, L. Vandepitte, D. Hobern, P. Schalk, R. E. DeWalt, K. Ma, J. Miller, T. Orrell, R. Aalbu, J. Abbott, R. Adlard, C. Aedo, et al., Catalogue of Life Checklist (1.1.24.106); https://doi.org/10.48580/dg4lg-49xk
[MBE] Moths and Butteflies of Europe
Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MPG]
North American Moth Photographers Group
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Michael Borsch]
Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes
[Michel Tarrier]
Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org
[NBC] Netherlands Centre for Biodiversity Naturalis
NBC naturalis
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Opler] Opler & Warren, 2003
Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[Pertti Pakkanen]
e-mail: Pertti Pakkanen; Butterfly and moth observations from Finland
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Tero Piirainen] Tampere, Finland
e-mail: Tero Piirainen
[Vitaly Charny]
Birmingham, Alabama, USA
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[ZOBODAT] Oberösterreichische Landesmuseum
OÖ. Landesmuseum; ZOBODAT - Literatur
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[²] This may require parentheses or not (original genus not verified)

Some related literature:

Acerbi, 1802
Travels through Sweden, Finland and Lapland to North Cape, in the years 1798 and 1799 Travels North Cape 2 : 3-380, pl. 1-3
Austin, 1983
A new subspecies of Speyeria atlantis from the great basin of Nevada J. Lep. Soc. 37 (3) : 244-248
Austin, 1998
New subspecies of Nymphalidae (Lepidoptera) from Nevada and Arizona Syst. W. N. Am. Butts. (46) : 573-586
Barnes, 1897
Some new species and varieties of Lepidoptera from the Western U. S. Can. Ent. 29 (2) : 39-42
Barnes & Benjamin, 1926
Notes on diurnal Lepidoptera, with addition and corrections to the recent "List of Diurnal Lepidoptera" Bull. South. Calif. Acad. Sci. 25 (3) : 88-98
Barnes & McDunnough, 1913
New North American Lepidoptera with notes on described species Contr. nat. Hist. Lep. N. Am. 2 (3) : 93-165, pl. 1-9, I-IX
Behr, 1862
[Our California Argynnides] Proc. Cal. Acad. Sci. 2 : 172-177
Behr, 1863
On Californian Lepidoptera - No. III Proc. Calif. Acad. Sci. 3 (1) : 84-93
Behr, 1864
Notes on the Argynnides of California Proc. Ent. Soc. Philad. 3 (3) : 434-436
Boisduval, 1852
Lépidoptères de la Californie Ann. Soc. ent. Fr. (2) 10 (2) : 275-324
Boisduval, 1869
Lépidoptères de la Californie Ann. Soc. ent. Belg. 12 : 5-28,37-94
Bramson, 1890
Die Tagfalter (Rhopalocera) Europas und des Caucasus Tagfalter Eur. Caucasus : 1-148
Brown, 1983
The type of Argynnis apacheana Skinner J. Lep. Soc. 37 (1) : 79-80
Chermock & Chermock, 1940
Some new diurnal Lepidoptera from the Riding Mountains and the Sand Ridge, Manitoba Can. Ent. 72 (4) : 81-83
Chermock & Frechin, 1947
A new Speyeria from Washington Pan-Pacific Ent. 23 (3) : 111-112
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2 Mon. Rhop. Sinesium : 1-854
Chou, Yuan, Yin, Zhang & Chen, 2002
New species, new subspecies, and new records of butterflies from China (VI) Entomotaxonomia 24 (1): 52-68
Clark, 1932
The butterflies of the District of Columbia and vicinity Bull. U.S. nat. Mus. 157 : 1-337
Cockerell, 1900
A New Variety of Argynnis Nitocris Ent. News 11 (10) : 622
Cockerell, 1909
Notes on Butterflies Ent. Rec. J. Var. 21 (7/8) : 186
Comstock, 1925
Studies in Pacific Coast Lepidoptera Bull. South. Calif. Acad. Sci. 24 (1): 3-4, (2): 35-37, pl. 2, (3): 62-68
Cramer, [1775]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 1 (1-7): 1-132, pl. 1-84 (1775), (8): 133-155, pl. 85-96 (1776)
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
de Nicéville, 1889
On new and little-known Butterflies from the Indian Region, with Revision of the Genus Plesioneura of Felder and of Authors J. Bombay nat. Hist. Soc. 4 (3) : 163-194, pl. A-B
dos Passos, 1935
Some butterflies of southern Newfoundland with descriptions of new subspecies Can. Ent. 67 (4) : 82-88
dos Passos & Grey, 1942
Two new North American subspecies of Argynnis, with some revisional notes Amer. Mus. Novit. 1214: 6pp, 1fig
dos Passos & Grey, 1945
A new species and some new subspecies of Speyeria Amer. Mus. Novit. 1297: 17pp, 30fgs
dos Passos & Grey, 1947
Systematic catalogue of Speyeria with designations of types and fixations of type localities Amer. Mus. Novit. 1370: 30pp
dos Passos & Grey, 1957
A new name for Argynnis lais Edwards J. N.Y. Ent. Soc. 63: 95-96
Drury, [1773]
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 1 : pl. 1-50 (1770), (1773) , : 1-123 (1837)
Dunford, 2009
Taxonomic overview of the greater fritillary genus Speyeria Scudder and the atlantis - hesperis species complexes, with species accounts, type images, and relevant literature (Lepidoptera: Nymphalidae) Insecta Mundi 90 : 1-74
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1862
Descriptions of certain Species of Diurnal Lepidoptera found within the limits of the United States and of British America. - No. [1]-3 Proc. Acad. nat. Sci. Philad. 13: 160-164 (1861), 14: 54-58, 221-226 (1862)
Edwards, 1862
Descriptions of certain species of Diurnal Lepidoptera found within the United States, figured in Doubleday's Genera but undescribed Proc. Ent. Soc. Philad. 1 (7) : 221-224
Edwards, 1864
Descriptions of certain species of Diurnal Lepidoptera, found within the limits of the United States and British America. (No. 1-3) Proc. ent. Soc. Philad. 2 (1): 14-22, pl. 1, 11 (1863), (2): 78-82 (1863), (4): 501-507 (1864)
Behr in Edwards, 1864; Edwards, 1864
Notes on the Argynnides of California Proc. ent. Soc. Philad. 3 : 434-436
Edwards, 1869
The Butterflies of North America. Vol. I Butts. N. Amer. 1: 218pp, 50pls
Edwards, 1890
The Butterflies of North America. Vol. III Butts N. Amer. 3: 431pp, 51pls
Edwards, 1869
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 2 (4) : 369-376
Edwards, 1870
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 3 (1) : 10-22
Edwards, 1870
Descriptions of new North American diurnal Lepidoptera Trans. amer. ent. Soc. 3 : 189-196
Edwards, 1872
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 4 : 61-70
Edwards, 1874
Descriptions of new species of diurnal Lepidoptera found in North America Trans. amer. ent. Soc. 5 (1): 13-19, (2): 103-111
Edwards, 1876
Description of new species of diurnal Lepidoptera found within the United States and British North America Trans. amer. ent. Soc. 5 : 202-208
Edwards, [1877]
New species of diurnal Lepidoptera Trans. amer. ent. Soc. 5 : 289-292
Edwards, 1877
Descriptions of new species of diurnal Lepidoptera found in North America Field & Forest 3 (5): 86-89, : 101-105
Edwards, 1878
Descriptions of new species of North American Lepidoptera Field & Forest 3 : 115-119, 142-144
Edwards, 1879
Descriptions of new species of butterflies collected by Mr. H. K. Morrison in Nevada, 1878; also, Remarks on some errors of synonymy and arrangement Can. Ent. 11 (3): 49-56, : 79-80
Edwards, 1879
Descriptions of new species of North American butterflies; also, Notes upon certain species Can. Ent. 11 (5) : 81-89
Edwards, 1881
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. Amer. Ent. Soc. 9 : 1-8
Edwards, 1883
Descriptions of new species of diurnal Lepidoptera found in British America and the United States Can. Ent. 15 (2) : 32-36
Edwards, 1883
Descriptions of new species of North American butterflies Can. Ent. 15 (11) : 209-211
Edwards, 1886
Description of new species of butterflies found in the United States Can. Ent. 18 (4) : 61-65
Edwards, 1892
Miscellaneous notes on butterflies, larvae, etc. Can. Ent. 24 (3): 49-56, (5): 105-111
H. Edwards, 1877
Pacific Coast Lepidoptera. - No. 22. Notes on some Diurnal Lepidoptera, with description of new varieties Proc. Calif. Acad. Sci. 7 (12) : 163-173
H. Edwards, 1877
Pacific Coast Lepidoptera. No. 26. - Description of two new forms of Diurnals Proc. Cal. Acad. Sc.: 1 (Dec. 17)
Ehrmann, 1917
Some New North American Butterflies Lepidopterist 1 (7) : 54-56
Eisner, 1942
Einige ergebnisse der Sichtung der Gattungen Melitaea und Argynnis im Rijksmuseum van Natuurlijke Historie, Leiden Zool. Meded. Leiden 24 (4) : 116-124
Elwes, 1889
A revision of he genus Argynnis Trans. ent. Soc. Lond. 1889 : 535-575
Evans, 1912
A list of Indian butterflies J. Bombay nat. Hist. Soc. 21 (2): 553-584, (3): 969-1008
Evans, 1924
The identification of Indian Butterflies (5-8) J. Bombay nat. Hist. Soc. 30 (1): 72-96, pl. 21-24, (2): 322-351, pl. 26-27, (3): 610-639, pl. 28, (4): 726-776, pl. 29
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus Syst. Ent. : 1-832
Fabricius, 1787
Mantissa insectorum sistens species nuper detectas adiectis synonymis, observationibus, descriptionibus, emendationibus Mantissa Insectorum 2 : 1-382
Fawcett, 1904
On some New and Little-known Butterflies, mainly from high elevations in the N.E. Himalayas Proc. zool. Soc. Lond. 2 (1) : 131-141, pl. 9
Fisher, 1858
Description of a New Species of ARGYNNIS Proc. Acad. Nat. Sci. Philad. 10 : 179-180
Fruhstorfer, 1903
Neue Nymphaliden aus Ostasien Dt. ent. Z. Iris 16 (1) : 22-42, pl. 1, f. 5-6 (early October)
Fruhstorfer, 1906
Rhopaloceren-Aberrationen Ent. Zs. 20 (3) : 17-18, f. 1-5 (15 April)
Fruhstorfer, 1907
Neue Argynnis aus der aglaja-Gruppe Int. ent. Zs. 1 (34) : 256-257 (16 November)
Fruhstorfer, 1908
Eine neue Argynnis-Rasse aus Spanien Ent. Zs. 22 (39) : 161 (26 December)
Fruhstorfer, 1912
6. Familie: Nymphalidae in Seitz, Gross-Schmett. Erde 9 : 453-456 (31 January, 1912), : 457-488 (18 March, 1912), : 489-496 (25 May, 1912), : 489-496 (10 May, 1912), : 497-504 (15 April, 1912), : 505-512 (10 May, 1912), : 513-528 (11 April, 1912), : 529-552 (20 April, 1912), : 553-592 (15 May, 1912), : 593-616 (25 September, 1912), : 617-632 (15 October, 1912), : 633-640 (25 October, 1912), : 641-656 (10 January, 1913), : 657-664 (12 January, 1913), : 665-680 (13 January, 1913), : 681-688 (25 January, 1913), : 689-696 (16 June, 1913), : 697-704 (5 August, 1913), : 705-712 (20 January, 1914), : 713-720 (20 November, 1913), : 721-744 (9 March, 1914), : 745-751 (30 July, 1915), : 753-760 (28 March, 1914), : 761-766 (14 April, 1914)
Gatrelle, 1998
Two new Nymphalidae from western North Carolina: new subspecies of Speyeria aphrodite and Phyciodes batesii Taxonomic Rep. 1 (3) : 1-8
Godman & Salvin, [1901]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-741 (?782), 3: pl. 1-112
Grey, 1989
Sundry Argynnine concepts revisited (Nymphalidae) J. Lep. Soc. 43 (1): 1-10
Grum-Grshimailo, 1890
Le Pamir et sa faune lépidoptérologique in Romanoff, Mém. Lép. 4 : 1-576, pl. A,1-21
Gunder, 1924
Several New Aberrant Lepidoptera (Rhopalocera) from California Ent. News 35 (5) : 153-158
Gunder, 1925
Several New Varieties of and Aberrant Lepidoptera (Rhopalocera) from California Ent. News 36 : 1-9, pl. 1
Gunder, 1927
New transition forms (Lepid., Rhopalocera) Can. Ent. 59 (12) : 281-287
Gunder, 1929
New butterflies and sundry notes (Lepidoptera, Rhopalocera) Bull. Brooklyn ent. Soc. 24 (5) : 325-332, pl. 30-31
Gunder, 1931
Some new Butterflies (Lepid., Rhopalocera) Bull. South. Calif. Acad. Sci. 30 (2) : 45-48
Gunder, 1932
New Rhopalocera Can. Ent. 64 (12) : 276-284
Gunder, 1933
Additional New Rhopalocera (Lepidoptera) Can. Ent. 65 (8) : 171-173
Gunder, 1934
Various new butterflies (Lepid., Rhopalocera) Can. Ent. 66 (6) : 125-131
Guppy & Kondla, 2002
The correct spellings and type localities of butterflies named from Mount Rainier, Washington, USA J. Lep. Soc. 56 (3): 180-131
Hammond & Dornfeld, 1983
A new subspecies of Speyeria egleis from the pumice region of central Oregon J. Lep. Soc. 37 (2) : 115-120
Hemming, 1933
Holarctic butterflies: miscellaneous notes on nomenclature Entomologist 66 (12) : 275-279
Hemming, 1942
On the correct name of the species commonly known as Argynnis aglaja (Linnaeus, 1758) (Lep. Nymphalidae) and matters incidental hereto Proc. R. ent. Soc. Lond. (B) 11 (11) : 155-160
[H&R]; Higgins & Riley, 1970
A Field Guide to the Butterflies of Britain and Europe
[NACL]; Hodges, 1983
Check List of the Lepidoptera of America North of Mexico Check List Lep. Am. N of Mexico
Holland, 1900
A Description of a Variety of Argynnis Nitocris from Chihuahua, Mexico Ent. News 11 (1) : 332-333
Holland, 1931
Notes on some American butterflies mainly relating to classification and nomenclature. Part 2 & 3 Ann. Carnegie Mus. 20 (1): 39-55 (1930), (2): 255-264 (1931)
Holland, 1988
A new subspecies of Speyeria atlantis (Nymphalidae) from south-central New Mexico Bull. Allyn Mus. 115: 1-9
Holland, 2010
A new subspecies of Argynnis nokomis from Sacramento Mountains of New Mexico J. Res. Lepid. 42 : 79-80
Holmgren, 1888
Aberrationer till Argynnis aglaja L och adippe L. Ent. Tidskr. 9 : 103-106
Jachontov, 1909
[Notice sur les Lèpidoptères Rhopalocères du Caucase] Revue Russe Ent. 8 : 282-292
Janson, 1877
Notes on Japanese Rhopalocera with the description of new species Cistula ent. 2 : 153-160
Kardakoff, 1928
Zur Kenntnis der Lepidopteren des Ussuri-Gebietes Ent. Mitt. 17 (4): 261-273, (8): 414-425
Kershaw, 1952
Two New Aberrations of Argynnis aglaia Linn. Ent. Rec. J. Var. 64 : 69-70, pl. 3
Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian) Eversmannia Suppl. 2 : 1-123
Kudrna, 1974
An annotated List of Japanese Butterflies Atalanta 5 : 92-120
Kuwayama, 1921
On some Aberrations of the Japanese Butterflies Dobuts. Z. 33 : 332-337
[NL4A]; Lamas, 2004
Checklist: Part 4A, Hesperioidea - Papilionoidea Atlas of Neotropical Lepidoptera
Lampa, 1885
Förteckning öfver Skandinaviens och Finlands Macrolepidoptera Ent. Tidskr. 6 (1) : 3-137
Lang & Han, 2009; Lang in Lang & Han, 2009
Study on some nymphalid butterflies from China Atalanta 40 (3/4) : 493-500
Lang in Lang & Wang, 2010
Study on some nymphalid butterflies from China - 2 Atalanta 41 : 221-228
[RFEL]; Leley, 2016
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera Cat. ins. Russian Far East. 2 : 1-812
Lempke, 1956
Catalogus der Nederlandse Macrolepidoptera (Vierde Supplement) Tijdschr. Ent. 99 (4) : 155-216
[BOW]; Lewis, 1974
Butterflies of the World
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Linnaeus, 1767
Systema Naturae per Regna tria Naturae, secundum Classes, Ordines, .... Editio Duocecima Reformata. Tom. 1. Part II. Syst. Nat. (Edn 12) 1 (2) : 533-1327, : Errata
Matsumura, 1908
Die Nymphaliden Japans Ent. Zs. 22 (39) : 157-158
Matsumura, 1911
Erster Beitrag zur Insekten Fauna von Sachalin J. Coll. Agr. Tohoku Imp. Univ. Sapporo 4 (1) : 1-145, pl. 1-2
Matsumura, 1928
New Butterflies especially from the Kuriles Insecta Matsumurana 2 (4) : 191-201
Matsumura, 1929
Some new butterflies from Korea received from Mr. T. Takamuku Insecta Matsumurana 3 (4) : 152-156
McCorkle & Hammond, 1988
Biology of Speyeria zerene hippolyta (Nymphalidae) in marine-modified environment J. Lep. Soc. 42 (3): 184-195
McDunnough, 1924
Some new Canadian argynnid races (Lepid.) Can. ent. 56 (2) : 42-43
McDunnough, 1927
The Lepidoptera of the Seton Lake Region, British Columbia Can. ent. 59 (7): 152-162, (8): 193-199, (9): 207-214, (10): 239-246, (11): 266-277
McDunnough, 1929
Notes on some diurnal Lepidoptera from Yellowstone Park and the adjacent regions of Montana Can. ent. 61 (5) : 105-107
McDunnough, 1935
A new race of Argynnis cybele from Nova Scotia Can. ent. 67 (1): 18-19
McDunnough, 1940
The argynnids of the Cariboo region of British Columbia (Lepid.) Can. ent. 72 (2) : 23-25
Ménétriés, 1832
Catalogue raisonne des objects de zoologie recuellis dans un voyage au Caucase et jusqu'aux frontières de la Perse Cat. Rais. : 1-271
Moeck, 1947
A New Subspecies of Speyeria atlantis (Edwards) from New Mexico (Lepidoptera: Nymphalidae) Ent. News 58 (3) : 73-75
Moeck, 1950
A New Subspecies of Speyeria atlantis (Edwards) from Nevada (Lepidoptera: Nymphalidae) Ent. News 61 (3) : 61-64
Moeck, 1957
Geographic variability in Speyeria. Comments, records and description of a new subspecies Milwaukee Ent. Soc., Special Paper: -
Moore, [1875]
Descriptions of new Asiatic Lepidoptera Proc. zool. Soc. Lond. 1874 (4) : 565-579, pl. 66-67
de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017
Interrelationships and diversification of Argynnis Fabricius and Speyeria Scudder butterflies Syst. Ent. 42 (4) : 635-649
Nakahara, 1926
Some Saghalien Butterflies: Notes and Descriptions Ins. Inscit. Mens. 14 : 45-48
Oberthür, 1909
Notes pour servir à établir la Faune Francaise et Algeriénne des Lépidoptères Étud. Lépid. Comp. 3 : 101-400, 405-415, pl. 11,16-20,23-26
Owen, 1893
Peculiar form of Argynnis erinna Edw Ent. News 4 (7) : 246
Pelham, 2008
A catalogue of the butterflies of the United States and Canada J. Res. Lepid. 40 : 1-652
Peschke, 1934
Neubeschreibungen und Ergänzungen der palaearktischen Lepidopterenfauna I. Int. ent. Zs. 28 : 430-432
Pflümer, 1879
Ein Beitrag zur Schmetterlingskunde Stettin Ent. Ztg 40 : 157-161
[NAB]; Pyle, 1981
Field Guide to North American Butterflies, National Audubon Society
Reakirt, 1866
Coloradian butteflies Proc. Ent. Soc. Philad. 6 (1) : 122-151
Rebel, [1909]
Fr. Berge's Schmetterlingsbuch nach dem gegenwärtigen Stande der Lepidopterologie in Berge, Schmetterlingsbuch (edn 9) : 1-509
Reiff, 1910
Argynnis cybele Fabr. Forma bartschi f. nov. Psyche 17 (6) : 252
Reuss, 1926
Systemischer uberlich der Dryadinae T. Rss mit einigen Neubeschreibungen Dt. ent. Zs. 1926 (1) : 65-70
Reuss, 1926
Über Funktion der Sexualarmaturen bei Lepidopteren (Rhop.) und die resultierende Weiterentwicklung meines versuchten natürlichen Systems der Dryadinae T. R. Dt. ent. Zs. 1926 (5) : 431-440
Riel, 1911
Argynnis aglaja L. aberr. hortensia Donzel Ann. Soc. Linn. Lyon (n.s.) 58 : 187-188
Rothschild, 1933
On a collection of Lepidoptera from Spanish Morocco Novit. Zool. 38 : 315-330
Röber, 1896
Neue Schmetterlinge aus dem cilicischen Taurus Ent. Nachr. 22 (6) : 81-84
de Sagarra, 1925
Anotacions a la Lepidopterologica ibèrica Butll. Inst. catal. Hist. nat. (2) 5 (1): 45-49, (9): 270-274
de Sagarra, 1926
Anotacions a la lepidopterologia Ibérica IV (1) Butll. Inst. catal. Hist. nat. (2) 6 (6-7) : 128-139
de Sagarra, 1930
Anotacions a la lepidopterologia Ibérica V (2). Formes noves de lepidòpters ibérics Butll. Inst. catal. Hist. nat. (2) 10 (7) : 110-118
Schuster von Forstner, 1928
Völlig schwarzer Perlmutterfalter Lepid. Rdsch. 2 (16) : 157-158
Scott, 1981
New Papilionoidea and Hesperioidea from North America Papilio (n.s.) 1 : 1-12
Kondla, Scott & Spomer in Scott, Kondla & Spomer, 1998; Scott, Kondla & Spomer, 1998; Spomer, Scott & Kondla in Scott, Kondla & Spomer, 1998
Speyeria hesperis and Speyeria atlantis are distinct species Papilio (N.S) 8 : 1-31
Scudder, 1872
A systematic Revision of some of the American Butterflies; with brief notes on those known to occur in Essex County, Mass. 4th Ann. Rep. Peabody Acad. Sci. (1871) : 24-83,errata
Scudder, 1875
Synonymic list of the butterflies of North Amercia, north of Mexico. (1) Nymphales Bull. Buffalo Soc. nat. Sci. 2 : 233-269
Scudder, 1889
The Butterflies of the Eastern United States and Canada, with special reference to New England Butts eastern U.S. Canada 1: 1-766, 2: 767-1774, 3: 1775-1958, pl. 1-89
[SPRK]; Seppänen, 1970
Suomen suurperhostoukkien ravintokasvit Animalia Fennica 14
Skinner, 1897
Notes on Rhopalocera, with descriptions of new species and varieties Can. Ent. 29 (6) : 154-156
Skinner, 1911
A new Argynnis and a new Parnassius (Lep.) Ent. News 22 (3) : 108
Skinner, 1917
New species of Lepidoptera Ent. News 28 (7) : 328-329
Skinner, 1918
Argynnis apacheana, a New Name (Lepid.) Ent. News 29 (2) : 67
Skinner, 1919
A New Species of Argynnis from Utah (Lepid., Rhop.) Ent. News 30 (8) : 216
Slaby, 1949
Faunistické paberky z Cech. (Lep.) - Faunistiques Lépidoptèlogogiques de la Boheme Acta Soc. ent. Cal. 45 (1-2) : 65-72
[EBW]; Smart, 1976
The Illustrated Encyclopedia of the Butterfly World Illust. Encyp. Butt. World: -
Staudinger in Staudinger & Rebel, 1901
Catalog der Lepidopteren des palaearktischen Faunengebiets. I. Famil. Papilionidae - Hepialidae Cat. Lep. palaearct. Faunengeb. 1 : 1-411
Stichel, 1901
Berichtigung und Ergänzung zu meinem Artikel über bemerkenswerthe Schmetterlings-Varietäten etc. Berl. ent. Z. 46 : 21-24
Strand, 1901
Beitrag zur Schmetterlingsfauna Norwegens Nyt Mag. Naturv. 39 (1) : 25-72
Strand, 1912
Ueber einige Nebenformen europäischer Großschmetterlinge Ent. Zs. 25 (49): 253-254, (50): 257-258
Strecker, 1878
Butterflies and Moths of North America. Complete Synonymical Catalogue of Macrolepidoptera ... Synon. Cat. Macroplep.: - : 1-281
[BMAT]; Tennent, 1996
The Butterflies of Morocco, Algeria and Tunisia : 1-217
Tutt, 1896
British Butterflies, being a popular hand-book for young stutends and collectors Brit. Butts : 1-476
[BRU2]; Tuzov, Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000
Guide to the Butterflies of Russia and adjacent territories: Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae Butts. Russia adj. terr. 2 : 1-580
Verity, 1914
Contributo allo studio della variazione nei Lepidotteri Boll. Soc. ent. ital. 45 : 203-238, pl. 1
Verity, 1919
Seasonal polymorphism and races of some European Grypocera and Rhopalocera Ent. Rec. J. Var. 31 : 26-31,43-48,87-89,121-129,178-184,193-201
Verity, 1922
Seasonal polymorphism and races of some European Grypocera and Rhopalocera Ent. Rec. J. Var. 34 : 12-15, 68-73, 89-93, 124-142
Verity, 1935
Notes on the geographical variations of Argynnis aglaja, L. Entomologist 68 : 189-191, : 200-204
Wileman, 1911
Aberrations of three Japanese butterflies Entomologist 44 : 55-56
Williams, 2002
Recognition of western populations of Speyeria idalia (Nymphalidae) as new subspecies J. Lep. Soc. 55 (4) : 144-149
Wolcott, 1916
Description of two Hitherto Undescribed Aberrations, the one of Argynnis alcestis, the of the Chlorippe celtis (Lep.) Ent. News, 27 (3) : 97-99, pl. 4
Wright, 1905
The Butterflies of the West Coast of the United States Butts. West Coast US : 3-257
Yakovlev, 2012
Checklist of Butterflies (Papilionoidea) of the Mongolian Altai Mountains, including descriptions of new taxa Nota lepid. 35 (1) : 51-96
Zhang, Cong, Shen, Opler & Grishin, 2020
Genomic evidence suggests further changes of butterfly names Taxonomic Rep. 8 (7) : 1-41

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Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.