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Speyeria Scudder, 1872
Speyeria Scudder, 1872; 4th Ann. Rep. Peabody Acad. Sci. (1871): 44, TS: Papilio idalia Drury
= Argynnis ; [Wahlberg]
Semnopsyche Scudder, 1875; Bull. Buffalo Soc. nat. Sci. 2 : 238, 258, TS: Papilio diana Cramer
= ; [NL4A, 261]
Mesoacidalia Reuss, 1926; Dt. ent. Z. 1926 (1) : 69, TS: Papilio aglaja Linnaeus
Proacidalia Reuss, 1926; Dt. ent. Z. 1926 (1) : 69, TS: Argynnis clara Blanchard
Neoacidalia Reuss, 1926; Dt. ent. Z. 1926 (1) : 69, TS: Papilio cybele Fabricius
Speyeria (Argynnini) ; [NL4A, 261]


[sp-3.jpg] 700x734 (~72Kb) USA: nr Crystal Mountain (47°13'26N 120°40'11W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela [sp-4.jpg] 600x503 (~56Kb) USA: Cape Blanco, (42°49'57N 124°32'50W ±1km), Curry Co. OR, 2.8.2001, Photo © Markku Savela

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.



fiSpeyeria aglaja (Linnaeus, 1758)
EU, Morocco, Iran - Siberia, AC, China, Korea, Japan. See [About maps] Orvokkihopeatäplä Dark Green Fritillary Stor pärlemorfjäril Dukatpärlefjäril Großer Perlmutterfalter Le Grand Nacré Lunares de Piata Dostojka aglaja
Papilio aglaja Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 481, TL: Sweden
Papilio pasilhoe Linnaeus, 1767; Syst. Nat. (Edn 12) 1 (2): 755 (repl. for Papilio aglaja Linnaeus)
Papilio charlotta Haworth, 1802; Prodr. Lep. Brit.: 3
?Argynnis aglaia v. basalis Matsumura, 1908; Ent. Zs. 22 (39) : 158, TL: Hokkaido
?Argynnis aglaja methana Fruhstorfer, 1908; Ent. Zs. 22 (39) : 161, TL: Sierra de Guadarrama, Castilia
Argynnis aglaia[sic] myonia ab. isshikii Matsumura, 1928; Insecta Matsumurana 2 (4) : 193, TL: Hokkaido
Argynnis aglaja, NSG Voucher Specimen [Wahlberg]
Argynnis aglaja, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Argynnis aglaja, Svenska fjärilar [SVF]
Argynnis aglaja, Norges Sommerfugler [NORLEP]
Argynnis aglaja, Moths and Butterflies of Europe [MBE]
Argynnis aglaja, Europaische Schmetterlinge [Mario Maier]
Argynnis aglaja, Captain's European Butterfly Guide [Simon Coombes]
Mesoacidalia aglaja ; [BOW: pl. 3, f. 13-14]
Mesoacidalia aglaia[sic, recte aglaja] ; [EBW]
Speyeria aglaja ; [MRS, 473]
Argynnis aglaja ; [BRU2: 36, pl. 23, f. 1-15]; [Otakar Kudrna]
[aglaja-1u.jpg] 500x363 (~61Kb) upperside West down, Devo, UK 14/06/95, Photo © Simon Coombes [aglaja-1v.jpg] 500x389 (~50Kb) underside West down, Devo, UK 14/06/95, Photo © Simon Coombes [aglaja-2u.jpg] 700x544 (~41Kb) upperside Finland, Kangasala, Ruutana, 682:34, 8/1997, Photo © Tero Piirainen
[aglaja-3.jpg] 780x696 (~49Kb) FINLAND: Ab: Suomusjärvi, 669:31, 29.6.1997, Photo © Markku Savela [aglaja-4o.jpg] 500x667 (~41Kb) ova Finland, Sipoo 670:39, 22.7.1980, Photo © Pertti Pakkanen [aglaja-5l.jpg] 550x371 (~71Kb) larva Central Europe 1990-2000, Photo © Michael Borsch
[aglaja-5p.jpg] 550x371 (~80Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch [aglaja-5u.jpg] 597x400 (~59Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch [aglaja-6fau.jpg] 777x565 (~100Kb) upperside female Russia, Moscow area, 29.7.2008, Photo © D. Smirnov
[aglaja-6fbu.jpg] 551x607 (~112Kb) upperside female Russia, Moscow area, 29.7.2008, Photo © D. Smirnov

Larva on Viola verecunda, Polygonum suffultum [MRS], Viola bicolor, Bistorta major [BRU2, 36]


SEU, CEU, Caucasus, Altai, Sayan, W.Siberia, S.Siberia Argynnnis aglaja aglaja
auxo Jachontov, 1908
Argynnnis aglaja aglaja ; [BRU2, 36]
S. a. borealis (Strand, 1901)
EU, Siberia, Far East, Kamchatka Argynnis aglaja borealis Strand, 1901
Argynnis aglaja borealis ; [BRU2, 36]
S. a. lyauteyi (Oberthür, 1920)
Morocco (Middle Atlas) Argynnis lyauteyi Oberthür, 1920; Étud. Lépid. Comp. 17 : 48, pl. D, TL: Azrou (Morocco)
Argynnis lyauteyi, Moths and Butterflies of Europe [MBE]
Mesoacidalia liauteyi[sic, recte lyauteyi] ; [EBW, (name only)]
Mesoacidalia aglaja lyauteyi ; [BMAT: 50, pl. 15, f. 10-13]
Argynnis lyauteyi lyauteyi ;
[lyauteyi-1u.jpg] 435x600 (~32Kb) upperside Morocco, Ifrane, Moyen Atlas, Photo © Michel Tarrier [lyauteyi-2v.jpg] 600x435 (~33Kb) underside Morocco, Ifrane, Moyen Atlas, Photo © Michel Tarrier

S. a. excelsior (Rothschild, 1933)
Morocco (Rif Mts.) Argynnis aglaia excelsior Rothschild, 1933; Novit. Zool. 38 : 317-318, TL: Ketama (Morocco)
Mesoacidalia aglaja excelsior ; [BMAT: 50, pl. 15, f. 14-17]
Argynnis lyauteyi excelsior ;
[excelsior-1u.jpg] 600x435 (~46Kb) upperside Morocco, Djebel Tisirène, Rif, Photo © Michel Tarrier [excelsior-2v.jpg] 600x435 (~29Kb) underside Morocco, Djebel Tisirène, Rif, Photo © Michel Tarrier

S. a. ottomana (Röber, 1896)
Armenia, Talys, Kopet-Dagh Argynnis aglaja ottomana Röber, 1896; Ent. Nachr. 22 (6) : 82
Argynnis aglaja ottomana ; [BRU2, 36]
S. a. gigasvitatha (Verity, 1935)
Tian-Shan, Ghissar, Darvaz, Alai, S.Altai Argynnis aglaja gigasvitatha Verity, 1935; Ent. Rec. 68[?]: 202
Argynnis aglaja alaicola Verity, 1935; Ent. Rec. 68[?]: 202
Argynnis aglaja gigasvitatha ; [BRU2, 36]
S. a. vitatha (Moore, [1875])
Pamirs Argynnis vitatha Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4) : 568, TL: Cashmere
?Argynnis aglaja cannelata Peschke, 1934; Int. ent. Z. 28: 431, TL: Kashmir
Mesoacidalia vitatha ; [EBW, (name only)]
Speyeria aglaja vitatha ; [MRS, 473]
Argynnis aglaja vitatha ; [BRU2, 36]
S. a. clavimacula (Matsumura, 1929)
S.Ussuri Argynnis aglaia[sic] clavimacula Matsumura, 1929; Insecta Matsumurana 3 (4) : 154, TL: Korea
Speyeria aglaja clavimacula ; [MRS, 473]
Argynnis aglaja clavimacula ; [BRU2, 36]
S. a. kenteana (Stichel, 1901)
Transbaikalia, N.Ussuri, Amur Argynnis aglaja graeseri Kardakov, 1928; Entomol. Mitt. 17 (4): 270, TL: Ussuri
Argynnis aglaja kenteana ; [BRU2, 36]
S. a. tonnai (Matsumura, 1928)
Sakhalin Argynnis aglaja sachalinensis Matsumura, 1911; J. Coll. Agr. Tohoku Imp. Univ. Sapporo 4 (1) : 43 (preocc.)
Argynnis aglaia[sic] tonnai ab. otaniana Matsumura, 1928; Insecta Matsumurana 2 (4) : 192, TL: Saghalien
Argynnis aglaja tonnai ; [BRU2, 36]
S. a. matsumurai (Nakahara, 1926)
Kuriles Argynnis aglaja matsumurai Nakahara, 1926; Ins. Insec. Mens 14: 48
Argynnis aglaia[sic] chishimensis Matsumura, 1928; Insecta Matsumurana 2 (4) : 192
Argynnis aglaja matsumurai ; [BRU2, 36]
S. a. bessa (Fruhstorfer, 1907)
Speyeria aglaja bessa ; Huang, 2001

Speyeria alexandra (Ménétriés, 1832)
N.Iran, W.Kopet-Dagh, Asia Minor. See [About maps] Argynnis alexandra Ménétriés, 1832; Cat. Rais.: 246, TL: [Zuvand Plateau, Talysh Mts., Azerbaijan]
Argynnis alexandra, NSG Voucher Specimen [Wahlberg]
Mesoacidalia alexandra ; [BOW: pl. 198, f. 5]
Argynnis alexandra ; [BRU2: 36, pl. 22, f. 7-9]

Larva on Viola, V. canina, V. riviniana, V. tricolor [SPRK]


Speyeria clara (Blanchard, [1844])
Himalayas, Tibet, Kashmir, Sikkim. See [About maps] Argynnis clara Blanchard, [1844]; in Jacquemont, Voy. Inde 4 (Zool.): 20, pl. 2, f. 2-3
Argynnis clara manis Fruhstorfer, 1903; Dt. ent. Z. Iris 16 (1) : 41, TL: Sikkim
Argynnis clara ; de Nicéville, 1889, J. Bombay nat. Hist. Soc. 4 (3) : 165, pl. A, f. 6 ♀

 

Speyeria claudia Fawcett²
Himalayas. See [About maps] Mesoacidalia claudia ; [BOW: pl. 196, f. 10]

 

Speyeria diana (Cramer, [1777])
SE.USA. See [About maps] Diana
Papilio diana Cramer, [1777]; Uitl. Kapellen 2 (9-16): 4, 148, pl. 98, f. D, E
Speyeria diana, Butterflies of America [BOA]
Speyeria diana, Georgia Lepidoptera (USA) [GALEPS]
Semnopsyche diana ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 12
Speyeria diana ; [NACL, #4449]; [EBW]; [BOW: pl. 15, f. 15-16]; [Opler]
[diana-1a.jpg] 940x735 (~121Kb) USA: Alabama, 4.6.2005, Photo © Vitaly Charny [diana-1b.jpg] 1078x876 (~155Kb) USA: Alabama, 4.6.2005, Photo © Vitaly Charny [diana-1u.jpg] 1203x883 (~192Kb) upperside USA: Alabama, 4.6.2005, Photo © Vitaly Charny
[diana-1v.jpg] 1321x1137 (~214Kb) underside USA: Alabama, 4.6.2005, Photo © Vitaly Charny [diana-1x.jpg] 1267x989 (~137Kb) USA: Alabama, 4.6.2005, Photo © Vitaly Charny

Larva on Viola [NAB]



Speyeria cybele (Fabricius, 1775)
USA. See [About maps] Great Spangled Fritillary
Papilio cybele Fabricius, 1775; Syst. Ent.: 516
Papilio daphnis Cramer, 1775
baal (Strecker, 1878)
Speyeria cybele, E.H. Strickland Ent. Mus, U of [ALBERTA]
Speyeria cybele cybele, Butterflies of America [BOA]
Argynnis cybele, NSG Voucher Specimen [Wahlberg]
Speyeria cybele, Butterflies and Moths of North America [BAMONA]
Argynnis cybele ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria cybele ; [NACL, #4450]; [EBW]; [BOW: pl. 15, f. 14, 17]; [Opler]
[cybele-1u.jpg] 1024x761 (~81Kb) upperside USA: Taneum Creek, Wenatchee National Forest, (47°06'24N 120°54'44W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela [cybele-1v.jpg] 700x632 (~45Kb) underside USA: Taneum Creek, Wenatchee National Forest, (47°06'24N 120°54'44W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela [cybele-2mu.jpg] 1000x790 (~91Kb) upperside male USA: nr Crystal Mountain / FR9726 (47°14'35N 120°40'41W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela
[cybele-2mv.jpg] 800x639 (~62Kb) underside male USA: nr Crystal Mountain / FR9726 (47°14'35N 120°40'41W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela [cybele-3f.jpg] 700x986 (~93Kb) female USA: Mima Mounds Natural Area Preserve, Thurston Co., WA, 13.8.2001, Photo © Markku Savela [cybele-4u.jpg] 1300x909 (~199Kb) upperside USA: Alabama, 26.5.2001, Photo © Vitaly Charny
[cybele-4v.jpg] 1300x1065 (~179Kb) underside USA: Alabama, 26.5.2001, Photo © Vitaly Charny

Larva on Viola, V. rotundifolia [NAB]


S. c. novascotiae (McDunnough, 1935)
Nova Scotia Argynnis cybele novascotiae McDunnough, 1935; Can. ent. 67: (18-19), TL: Nova Scotia
Speyeria cybele novascotiae ; [NACL, #4450a]
S. c. krautwurmi (Holland, 1931)
Speyeria cybele krautwurmi, Butterflies of America [BOA]
Speyeria cybele krautwurmi ; [NACL, #4450b]
S. c. pseudocarpenteri (F. & R. Chermock, 1940)
Speyeria cybele pseudocarpenteri ; [NACL, #4450c]
S. c. carpenterii (Edwards, 1876)
cybele carpenterii (Edwards, 1876); Trans. amer. ent. Soc. 5 : 204
Argynnis cybele carpenterii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria cybele carpenterii ; [NACL, #4450d]
S. c. charlottii (Barnes, 1897)
Speyeria cybele charlottii, Butterflies of America [BOA]
Speyeria cybele charlottii ; [NACL, #4450e]
S. c. letona dos Passos & Grey, 1945
Speyeria cybele letona dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: City Creek Canyon, Salt Lake City, Utah
Speyeria cybele letona ; [NACL, #4450f]
S. c. pugetensis Chermock & Frechin, 1947
Washington cybele pugetensis Chermock & Frechin, 1947; Pan-Pacif. Ent. 23 (3): 111
Speyeria cybele pugetensis, Butterflies of America [BOA]
Speyeria cybele pugetensis ; [NACL, #4450g]
S. c. leto (Behr, 1862)
California, Alberta Argynnis leto Behr, 1862:REF#: 173; California
letis (Wright)
valesinoidesalba (Reuss, 1926)
lethe (Gunder, 1934)
Speyeria cybele leto, Butterflies of America [BOA]
Argynnis leto ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria cybele leto ; [NACL, #4450h]

Speyeria aphrodite (Fabricius, 1787)
Canada - Kansas. See [About maps] Aphrodite Fritillary
Papilio aphrodite Fabricius, 1787; Mantissa Insectorum 2: 62
?daphnis Martyn
= ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
baartschi (Reiff, 1910)
bakeri (Clark, 1932)
Speyeria aphrodite, E.H. Strickland Ent. Mus, U of [ALBERTA]
Speyeria aphrodite, Butterflies of America [BOA]
Argynnis aphrodite, NSG Voucher Specimen [Wahlberg]
Argynnis aphrodite ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria aphrodite ; [NACL, #4451]; [BOW: pl. 15, f. 5-6]; [Opler]

Larva on Viola, V. lanceolata, V. fimbriatula, V. nuttallii, V. primulifolia [NAB]

S. a. winni (Gunder, 1932)
Speyeria aphrodite winni, Butterflies of America [BOA]
Speyeria aphrodite winni ; [NACL, #4451a]
S. a. alcestis (Edwards, 1876)
aphrodite alcestis (Edwards, 1876); Trans. amer. ent. Soc. 5 : 289 (?[1877])
suffusa (Wolcott, 1916); Ent. News, 27 (3): (97-99)
Speyeria aphrodite alcestis, Butterflies of America [BOA]
Argynnis aphrodite alcestis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria aphrodite alcestis ; [NACL, #4451b]
S. a. manitoba (F. & R. Chermock, 1940)
mayae (Gunder, 1932); (ab.)
Speyeria mayae dos Passos & Grey, 1947
Speyeria aphrodite manitoba, Butterflies of America [BOA]
Speyeria aphrodite manitoba ; [NACL, #4451c]
S. a. whitehousei (Gunder, 1932)
Speyeria aphrodite whitehousei, Butterflies of America [BOA]
Speyeria aphrodite whitehousei ; [NACL, #4451d]
S. a. columbia (H. Edwards, 1877)
aphrodite columbia (H. Edwards, 1877); Proc. Cal. Acad. Sc.: ?
Speyeria aphrodite columbia, Butterflies of America [BOA]
Argynnis columbia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria aphrodite columbia ; [NACL, #4451e]
S. a. ethene (Hemming, 1933)
Argynnis cypris Edwards, 1886; (preocc. by Meigen, 1828)
Speyeria aphrodite ethene, Butterflies of America [BOA]
Speyeria aphrodite ethene ; [NACL, #4451f]
S. a. byblis (Barnes & Benjamin, 1926)
Speyeria aphrodite byblis, Butterflies of America [BOA]
Speyeria aphrodite byblis ; [NACL, #4451g]

Speyeria idalia (Drury, [1773])
E.USA. See [About maps] Regal Fritillary
Papilio idalia Drury, [1773]; Illust. Nat. Hist. Exot. Insects 1: index, : 25, pl. 13, f. 1-3
astarte (Fisher, 1858); (preocc. by Doubleday, 1847)
astaroth (Fisher, 1859)
infumata (Oberthür, 1912)
dolli (Gunder, 1927)
pallida (Eisner, 1942)
Speyeria idalia idalia, Butterflies of America [BOA]
Speyeria idalia, Butterflies and Moths of North America [BAMONA]
Speyeria idalia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13; [NACL, #4452]; [EBW]; [BOW: pl. 16, f. 2]; [Opler]

Larva on Viola [NAB]

S. i. occidentalis Williams, 2002
Speyeria idalia occidentalis Williams, 2002; J. Lep. Soc. 55 (4): 146, TL: 1 mile SE of Crete, Illinois
Speyeria idalia occidentalis, Butterflies of America [BOA]

Speyeria nokomis (Edwards, 1862)
Utah, Arizona. See [About maps] Nokomis Fritillary
Argynnis nokomis Edwards, 1862; Proc. Acad. nat. Sci. Philad. 14: 221, TL: Mt. Sneffels, Ouray Co., Colorado
Argynnis nitocris var. nigrocaerulea Cockerell, 1900; Ent. News 11 (10) : 622, TL: Beulah, Sapello Canon, New Mexico
rufescens (Cockerell, 1909)
valesinoidesalba (Reuss, 1926); (preocc by Reuss, 1926)
Speyeria nokomis nokomis, Butterflies of America [BOA]
Speyeria nokomis, Butterflies and Moths of North America [BAMONA]
Argynnis nokomis ; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 674; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Argynnis nitocris nigrocaerulea ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria nokomis ; [NACL, #4453]; [EBW]; [BOW: pl. 16, f. 8]; [Opler]; [NL4A, #2289]

Larva on (eggs near) Viola nephrophylla? [NAB]

S. n. nitocris (Edwards, 1874)
Arizona Argynnis nitocris Edwards, 1874; Trans. amer. ent. Soc. 5 (1): 15
Speyeria nokomis nitocris, Butterflies of America [BOA]
nitrocris[sic, recte nitocris] ; dos Passos, 1964
Argynnis nitocris ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria nokomis nitocris ; [NACL, #4453a]
Speyeria nitocris ; [BOW: pl. 16, f. 7]
S. n. coerulescens (Holland, 1900)
Mexico Argynnis nokomis var. coerulescens Holland, 1900; Ent. News 11 (1) : 332, TL: Mexico, Chihuahua
Speyeria nokomis coerulescens, Listing of Mexican Butterflies [ILMB]
Speyeria nokomis coerulescens, Butterflies of America [BOA]
Speyeria nokomis coerulescens, Butterflies of Southeastern Arizon [Bruce Walsh]
caerulescens[sic, recte coerulescens] ; McDunnough, 1938
Argynnis nitocris var. coerulescens ; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 675, 3: pl. 112, f. 15-18
Argynnis nitocris caerulescens[sic, recte coerulescens] ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria nokomis coerulescens ; [NACL, #4453b]; [NL4A, #2289a]
S. n. apacheana (Skinner, 1918)
Argynnis apacheana Skinner, 1918; Ent. News 29 (2) : 67, TL: near Independence, Inyo Co., California
nokomis Edwards, 1873
hermosa (Comstock, 1925)
Speyeria nokomis apacheana, Butterflies of America [BOA]
apacheana ; Brown, 1983, J. Lep. Soc. 37 (1): 79-80
Speyeria nokomis apacheana ; [NACL, #4453c]
S. n. wenona dos Passos & Grey, 1945
Mexico Speyeria wenona dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Cerro Potosi, Municipio de Galeana, Nuevo Leon, Mexico, 12000ft
Speyeria nokomis wenona ; [NL4A, #2289c]
S. n. melaena Mooser & Garcia, 1980
Mexico nokomis melaena Mooser & Garcia, 1980; , TL: Mexico, Aguascalienes
Speyeria nokomis melaena ; [NL4A, #2289b]

Speyeria edwardsii (Reakirt, 1866)
Colorado, Utah, Nevada. See [About maps] Edward's Fritillary
Argynnis edwardsii Reakirt, 1866; Proc. Ent. Soc. Philad. 6 (1) : 137
aglaia Edwards, 1864
montana (Reuss, 1926)
edonis (Gunder, 1934)
Speyeria edwardsii, E.H. Strickland Ent. Mus, U of [ALBERTA]
Argynnis edwardsii, NSG Voucher Specimen [Wahlberg]
Speyeria edwardsii, Butterflies and Moths of North America [BAMONA]
Argynnis edwardsii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria edwardsii ; [NACL, #4454]; [EBW]; [BOW: pl. 15, f. 19]; [Opler]

Larva on Viola, V. nuttallii [NAB]


Speyeria coronis (Behr, 1864)
California. See [About maps] Coronis Fritillary
Argynnis coronis Behr, 1864; in Edwards, Proc. ent. Soc. Philad. 3 : 435
Argynnis californica Skinner, 1917; Ent. News 28 (7) : 328
Speyeria coronis coronis, Butterflies of America [BOA]
Argynnis coronis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria coronis ; [NACL, #4455]; [BOW: pl. 15, f. 13]; [Opler]; [NL4A, #2288]
[coronis-1vU.jpg] identification uncertain 600x834 (~64Kb) underside USA: FR 9705 x US Route 97, (47°18'38N 120°41'48W), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela [coronis-2U.jpg] identification uncertain 600x762 (~49Kb) USA: Table Mtn, Kittitas Co., WA, 30.7.1999, Photo © Markku Savela [coronis-3vU.jpg] identification uncertain 600x619 (~54Kb) underside USA: FR9708, Table Mtn, Kittitas Co., WA, 19.7.1999, Photo © Markku Savela

Larva on Viola [NAB]


S. c. hennei (Gunder, 1934)
Speyeria coronis hennei, Butterflies of America [BOA]
Speyeria coronis hennei ; [NACL, #4455a]
S. c. semiramis (Edwards, 1886)
WNA, California Argynnis semiramis Edwards, 1886; Can. Ent. 18 (4) : 61, TL: California
Argynnis semiramis tr.f. bernardensis Gunder, 1933; , TL: California
= ; [NL4A, #2288a]
Speyeria coronis semiramis, Listing of Mexican Butterflies [ILMB]
Speyeria coronis semiramis, Butterflies of America [BOA]
Argynnis semiramis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria semiramis ; [BOW: pl. 16, f. 10]
Speyeria coronis semiramis ; [NACL, #4455b]; [NL4A, #2288a]
S. c. simaetha dos Passos & Grey, 1945
Speyeria coronis simaetha dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Black Canyon, Cascade Mts., nr Brewster, Washington
Speyeria coronis simaetha ; [NACL, #4455d]
S. c. snyderi (Skinner, 1897)
Utah coronis snyderi (Skinner, 1897); Can. Ent. 29 (6) : 154
Argynnis snyderi ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria coronis snyderi ; [NACL, #4455e]
S. c. halcyone (Edwards, 1869)
Colorado coronis halcyone (Edwards, 1869); Butts. N. Amer. 1: 83
Argynnis halcyone ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria halcyone ; [BOW: pl. 15, f. 20]
Speyeria coronis halcyone ; [NACL, #4455f]

Speyeria zerene (Boisduval, 1852)
California. See [About maps] Zerene Fritillary
zerene (Boisduval, 1852); Ann. Soc. ent. Fr. (2) 10 : 303, TL: "Yosemite"? or Agua Fria, W of Mariposa
Argynnis monticola Behr, 1862; Proc. Cal. Acad. Sci. 2 : 175 (?[1863])
Speyeria zerene, E.H. Strickland Ent. Mus, U of [ALBERTA]
Speyeria zerene zerene, Butterflies of America [BOA]
Argynnis zerene, NSG Voucher Specimen [Wahlberg]
Argynnis monticola ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Argynnis zerene ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria zerene ; [NACL, #4456]; [EBW]; [BOW: pl. 16, f. 9, 11]; [Opler]
[zerene-1vU.jpg] identification uncertain 700x881 (~88Kb) underside USA: FR 9712 nr. Liberty, Lion Gulch, (47°15'47N 120°40'15W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela [zerene-2U.jpg] identification uncertain 700x904 (~73Kb) USA: FR 9705 nr. Liberty and Lion Gulch, (47°17'49N 120°39'58W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela [zerene-3vU.jpg] identification uncertain 600x787 (~65Kb) underside USA: FR 9705 nr. US Route 97, (47°18'15N 120°41'24W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

Larva on Viola, V. adunca, Viola cuneata, V. lobata [NAB]


S. z. conchyliata (Comstock, 1925)
E.Oregon (E slope of Oregon Cascade) shastaensis (Comstock, 1925)
Speyeria zerene conchyliata ; [NACL, #4456a]
Speyeria zerene conchyliatus ;
S. z. gloriosa Moeck, 1957
Speyeria zerene gloriosa ; [NACL, #4456b]
S. z. sordida (Wright, 1905)
Speyeria zerene sordida ; [NACL, #4456c]
S. z. malcomi (Comstock, 1920)
sineargentatus (Gunder, 1927)
Speyeria zerene malcomi ; [NACL, #4456d]
S. z. hippolyta (Edwards, 1879)
coastal (California, W.Oregon, W.Washington) Argynnis hippolyta Edwards, 1879; Can. Ent. 11 (5) : 81
Speyeria zerene hippolyta, Butterflies of America [BOA]
Argynnis hippolyta ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria zerene hippolyta ; [NACL, #4456f]; McCorkle & Hammond, 1988, J. Lep. Soc. 42 (3): 184-195
Speyeria hippolyta ; [BOW: pl. 15, f. 21]

Larva on Viola adunca

S. z. bremnerii (Edwards, 1872)
inland (Vancouver Island - W.Washington - W.Oregon) zerene bremnerii (Edwards, 1872); Trans. amer. ent. Soc. 4 : 63
Argynnis bremnerii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria zerene bremnerii ; [NACL, #4456i]
S. z. behrensii (Edwards, 1869)
zerene behrensii (Edwards, 1869); Trans. amer. ent. Soc. 2 : 370
Argynnis behrensii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria zerene behrensii ; [NACL, #4456g]
S. z. myrtleae dos Passos & Grey, 1945
Speyeria zerene myrtleae dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: San Mateo, California
Speyeria zerene myrtleae ; [NACL, #4456h]
S. z. gunderi (Comstock, 1915)
Speyeria zerene cynna dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Humboldt Nat. Forest, Ruby Valley, Elko Co., NV
= ; Grey, 1989, J. Lep. Soc. 43 (1): 3
Speyeria zerene gunderi, Butterflies of America [BOA]
Speyeria zerene cynna ; [NACL, #4456o]
Speyeria coronis gunderi ; [NACL, #4455c]
Speyeria zerene gunderi ; Grey, 1989, J. Lep. Soc. 43 (1): 3
S. z. picta (McDunnough, 1924)
Speyeria zerene picta ; [NACL, #4456j]
S. z. garretti (Gunder, 1932)
Speyeria zerene garretti ; [NACL, #4456k]
S. z. sinope dos Passos & Grey, 1945
Speyeria zerene sinope dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Estes Park area, Rocky Mt. National Park, Colorado
Speyeria zerene sinope ; [NACL, #4456l]
S. z. platina (Skinner, 1897)
Utah, Idaho zerene platina (Skinner, 1897); Can. Ent. 29 (6) : 154
zerene pfoutsi (Gunder, 1933)
= ; Grey, 1989, J. Lep. Soc. 43 (1): 3
Argynnis platina ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria zerene pfoutsi ; [NACL, #4456n]
Speyeria zerene platina ; [NACL, #4456m]

Speyeria carolae (dos Passos & Grey, 1942)
Argynnis coronis carolae dos Passos & Grey, 1942; Amer. Mus. Novit. 1214: (1-6), TL: Charleston Park, Clark Co., NE
Speyeria carolae, Butterflies of America [BOA]
Speyeria carolae, Butterflies and Moths of North America [BAMONA]
Speyeria zerene carolae ; [NACL, #4456e]
Speyeria carolae ; [Opler]

Speyeria callippe (Boisduval, 1852)
California. See [About maps] Callippe Fritillary
Argynnis callippe Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 : 306
Speyeria callippe, E.H. Strickland Ent. Mus, U of [ALBERTA]
Speyeria callippe callippe, Butterflies of America [BOA]
Argynnis callippe, NSG Voucher Specimen [Wahlberg]
Speyeria callippe, Butterflies and Moths of North America [BAMONA]
Argynnis callippe ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria callippe ; [NACL, #4457]; [BOW: pl. 15, f. 11]; [Opler]; [NL4A, #2287]

Larva on Viola, Viola pedunculata, V. nuttallii [NAB]

S. c. comstocki (Gunder, 1925)
California Argynnis calippe[sic] var. comstocki Gunder, 1925; Ent. News 36 : 8, TL: California
Argynnis comstocki tr.f. creelmani Gunder, 1934; , TL: California
= ; [NL4A, #2287a]
Speyeria callippe comstocki, Listing of Mexican Butterflies [ILMB]
Speyeria callippe comstocki, Butterflies of America [BOA]
Speyeria callippe comstocki ; [NACL, #4457a]; [NL4A, #2287a]
S. c. liliana (H. Edwards, 1877)
Argynnis liliana H. Edwards, 1877; Proc. Calif. Acad. Sci. 7 : 170
Argynnis liliana var. baroni Edwards, 1881; Trans. Amer. Ent. Soc. 9 : 3
Speyeria callippe liliana, Butterflies of America [BOA]
Argynnis liliana ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Argynnis liliana baroni ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria callippe liliana ; [NACL, #4457b]
S. c. semivirida (McDunnough, 1924)
Speyeria callippe semivirida, Butterflies of America [BOA]
Speyeria callippe semivirida ; [NACL, #4457c]
S. c. elaine dos Passos & Grey, 1945
Speyeria callippe elaine dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Butte Falls, Oregon
Speyeria callippe elaine ; [NACL, #4457d]
S. c. rupestris (Behr, 1863)
Argynnis rupestris Behr, 1863; Proc. Calif. Acad. Sci. 3 (1) : 84
?irene Edwards
= Argynnis rupestris ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Argynnis rupestris ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria callippe rupestris ; [NACL, #4457e]
S. c. juba (Boisduval, 1869)
Argynnis inornata Edwards, 1872; Trans. amer. ent. Soc. 4 : 64
Speyeria callippe juba, Butterflies of America [BOA]
Argynnis inornata ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria callippe juba ; [NACL, #4457f]
S. c. sierra dos Passos & Grey, 1945
Speyeria callippe sierra dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Gold Lake, Sierra Co., California
= ?Speyeria callippe juba (Boisduval, 1869); Grey, 1989, J. Lep. Soc. 43 (1): 4
Speyeria callippe sierra ; [NACL, #4457h]
S. c. laura (Edwards, 1879)
Argynnis laura Edwards, 1879; Can. Ent. 11 (3): 49
Argynnis laura ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria callippe laura ; [NACL, #4457g]
S. c. nevadensis (Edwards, 1870)
Utah, Nevada, Wyoming callippe nevadensis (Edwards, 1870); Trans. amer. ent. Soc. 3 (1) : 14
Speyeria callippe nevadensis, Butterflies of America [BOA]
Argynnis nevadensis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria nevadensis ; [BOW: pl. 16, f. 6]
Speyeria callippe nevadensis ; [NACL, #4457i]
S. c. macaria (Edwards, 1877)
California, Nevada Speyeria callippe macaria, Butterflies of America [BOA]
Speyeria callippe macaria ; [NACL, #4457j]
Speyeria macaria ; [BOW: pl. 16, f. 5]
S. c. laurina (Wright, 1905)
Argynnis wrightii Wright, 1905
Speyeria callippe laurina, Butterflies of America [BOA]
Speyeria callippe laurina ; [NACL, #4457k]
S. c. harmonia dos Passos & Grey, 1945
Speyeria callippe harmonia dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Mt. Wheeler, Snake Range, nr. Utah border, NV, 8000ft
Speyeria callippe harmonia ; [NACL, #4457l]
S. c. meadii (Edwards, 1872)
callippe meadii (Edwards, 1872); Trans. amer. ent. Soc. 4 : 69
Argynnis nevadensis r. meadii f. gerhardi Gunder, 1927
Speyeria callippe meadii, Butterflies of America [BOA]
Argynnis nevadensis meadii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria callippe meadii ; [NACL, #4457m]
S. c. gallatini (McDunnough, 1929)
Speyeria callippe gallatini, Butterflies of America [BOA]
Speyeria callippe gallatini ; [NACL, #4457n]
S. c. calgariana (McDunnough, 1924)
Canada Speyeria callippe calgariana ; [NACL, #4457o]
Speyeria calgariana ; [BOW: pl. 15, f. 10]

Speyeria egleis (Behr, 1862)
-. See [About maps] Egleis Fritillary
Argynnis egleis Behr, 1862; Proc. Cal. Acad. Sci. 2 : 174
Argynnis egleis Boisduval, 1869; Ann. Soc. ent. Belg. 12 : 59
Argynnis montivaga Behr, 1863; Proc. Calif. Acad. Sci. 3 (1) : 84
Argynnis astarte Edwards, 1864; Proc. ent. Soc. Philad. 3 : 434 (preocc. by Doubleday, 1847)
?nenoquis Reakirt
= ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Argynnis montivaga ab. mammothi Gunder, 1924; Ent. News 35 (5) : 157, pl. 2, f. H, D
Argynnis montivaga tr. f. boharti Gunder, 1929; Bull. Brooklyn ent. Soc. 24 (5): 326, pl. 31, f. 5
Argynnis egleis Boisduval, 1869
Speyeria egleis, E.H. Strickland Ent. Mus, U of [ALBERTA]
Speyeria egleis egleis, Butterflies of America [BOA]
Speyeria egleis, Butterflies and Moths of North America [BAMONA]
Argynnis montivaga ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Argynnis egleis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria egleis ; [NACL, #4458]; [Opler]

Larva on Viola adunca [NAB]

S. e. clemencei (Comstock, 1925)
Speyeria egleis clemencei ; [NACL, #4458b]
S. e. atossa (Edwards, 1890)
California egleis atossa (Edwards, 1890); Butts N. Amer. 3: pl. 8
tejonica (Comstock, 1925)
Argynnis atossa ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria atossa ; [BOW: pl. 15, f. 8]
Speyeria egleis atossa ; [NACL, #4458c]
S. e. tehachapina (Comstock, 1920)
Speyeria egleis tehachapina, Butterflies of America [BOA]
Speyeria egleis tehachapina ; [NACL, #4458d]
S. e. moecki Hammond & Dornfeld, 1983
Oregon Speyeria egleis moecki Hammond & Dornfeld, 1983; J. Lep. Soc. 37 (2): 115, f. 2, TL: Skookum Mdw., Walker Rim, Klamath Co., Oregon
Speyeria egleis moecki, Butterflies of America [BOA]

Larva on Viola purpurea Hammond & Dornfeld, 1983, J. Lep. Soc. 37 (2): 120

S. e. oweni (Edwards, 1892)
S.Oregon - California oweni Edwards, 1892; Can. Ent. 24 (5): 105, TL: Mt. Shasta, Sikiyou Co., California
Speyeria egleis oweni, Butterflies of America [BOA]
Argynnys hippolyta oweni ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria egleis oweni ; [NACL, #4458e]; Hammond & Dornfeld, 1983, J. Lep. Soc. 37 (2): (note), f. 3
S. e. linda (dos Passos & Grey, 1942)
Argynnis utahensis linda dos Passos & Grey, 1942; Amer. Mus. Novit. 1214: (1-6), TL: Heyburn Peak, Sawtooth-Boise, ID, 9500-10000ft
Speyeria egleis linda ; [NACL, #4458f]
S. e. macdunnoughi (Gunder, 1932)
Speyeria egleis macdunnoughi, Butterflies of America [BOA]
Speyeria egleis macdunnoughi ; [NACL, #4458g]
S. e. albrighti (Gunder, 1932)
Speyeria egleis albrighti ; [NACL, #4458h]
S. e. utahensis (Skinner, 1919)
Speyeria egleis utahensis, Butterflies of America [BOA]
Speyeria egleis utahensis ; [NACL, #4458i]
S. e. toiyabe Howe, 1975
Speyeria egleis toiyabe ; [NACL, #4458j]
S. e. secreta dos Passos & Grey, 1945
Speyeria montiviga[sic] secreta dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Estes Park area, Rocky Mt. National Park, Colorado, 8000ft
Speyeria egleis secreta ; [NACL, #4458k]

Speyeria adiaste (Edwards, 1864)
California. See [About maps] Unsilverered Fritillary Adiaste Fritillary
?adiaste (Behr, 1862); Proc. Cal. Acad. Sci. 2 : 175
Argynnis adiante Boisduval, 1869; Ann. Soc. ent. Belg. 12 : 61
Argynnis adiaste Behr, 1862
Speyeria adiaste adiaste, Butterflies of America [BOA]
Speyeria adiaste, Butterflies and Moths of North America [BAMONA]
Speyeria adraste ; Kirby, 1871, (missp.)
Speyeria adianthe ; Barnes & McDunnough, 1917, (missp.)
Argynnis adiaste ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria egleis adiaste ; [NACL, #4458a]
Speyeria adiante ; [BOW: pl. 15, f. 4]
Speyeria adiaste ; [Opler]

Speyeria atlantis (Edwards, 1862)
ENA. See [About maps] Atlantis Fritillary
atlantis (Edwards, 1862); Proc. Acad. nat. Sci. Philad. 14: 54
Argynnis atlantis ab. chemo Scudder, 1889
Speyeria atlantis, E.H. Strickland Ent. Mus, U of [ALBERTA]
Speyeria atlantis atlantis, Butterflies of America [BOA]
Argynnis atlantis, NSG Voucher Specimen [Wahlberg]
Speyeria atlantis, Butterflies and Moths of North America [BAMONA]
Argynnis atlantis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria atlantis ; [NACL, #4459]; [BOW: pl. 15, f. 7]; [Opler]
[atlantis-1vU.jpg] identification uncertain 700x795 (~54Kb) underside USA: FR 9705 nr. US Route 97, (47°18'15N 120°41'24W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela [atlantis-2vU.jpg] identification uncertain 700x976 (~102Kb) underside USA: FR 9705 x US Route 97, (47°18'38N 120°41'48W), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

Larva on (eggs near) Viola adunca?, V. purpurea?, V. canadensis? [NAB]


S. a. canadensis (dos Passos, 1935)
Argynnis atlantis canadensis dos Passos, 1935; Can. Ent. 67: (82-88), TL: Doyles Station, Newfoundland
Speyeria atlantis canadensis, Butterflies of America [BOA]
Speyeria atlantis canadensis ; [NACL, #4459a]
S. a. hollandi (F. & R. Chermock, 1940)
Speyeria atlantis hollandi (F. & R. Chermock, 1940); [ NACL]
S. a. hesperis (Edwards, 1864)
Colorado, Utah, Montana atlantis hesperis (Edwards, 1864); Proc. ent. Soc. Philad. 2 (4): 502
Speyeria hesperis, E.H. Strickland Ent. Mus, U of [ALBERTA]
Speyeria hesperis, Butterflies and Moths of North America [BAMONA]
Argynnis hesperis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria hesperis ; [BOW: pl. 16, f. 1]; [Opler]
Speyeria atlantis hesperis ; [NACL, #4459c]
S. a. nikias (Ehrmann, 1917)
Speyeria atlantis nikias ; [NACL, #4459d]
S. a. dorothea Moeck, 1947
Speyeria atlantis dorothea ; [NACL, #4459e]
S. a. nausicaa (Edwards, 1874)
Argynnis aphrodite f. arizonensis Elwes, 1889
Speyeria atlantis nausicaa, Butterflies of Southeastern Arizon [Bruce Walsh]
Speyeria atlantis nausicaa ; [NACL, #4459f]
S. a. schellbachi Garth, 1949
shellbachi ; dos Passos, 1964, (missp.)
Speyeria atlantis schellbachi ; [NACL, #4459g]
S. a. chitone (Edwards, 1879)
Argynnis chitone Edwards, 1879; Can. Ent. 11 (5) : 82
Argynnis chitone ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria atlantis chitone ; [NACL, #4459h]
S. a. wasatchia dos Passos & Grey, 1945
Speyeria atlantis wasatchia dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Payson Canyon, Payson, Utah
Speyeria atlantis wasatchia ; [NACL, #4459i]
S. a. greyi (Moeck, 1950)
Speyeria atlantis greyi ; [NACL, #4459j]
S. a. tetonia dos Passos & Grey, 1945
Speyeria atlantis tetonia dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Teton Mts., WY
Speyeria atlantis tetonia ; [NACL, #4459k]
S. a. viola dos Passos & Grey, 1945
Speyeria atlantis viola dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Trail Creek, Sawtooth Mts., ID, 7400ft
Speyeria atlantis viola ; [NACL, #4459l]
S. a. dodgei (Gunder, 1931)
Speyeria atlantis dodgei ; [NACL, #4459m]
S. a. irene (Boisduval, 1869)
Argynnis egleis var. irene Boisduval, 1869; Ann. Soc. ent. Belg. 12 : 60
cottlei (Comstock, 1925)
Argynnis rupestris irene ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria atlantis irene ; [NACL, #4459n]
S. a. electa (Edwards, 1878)
Colorado, Montana Argynnis electa Edwards, 1878; Field & Forest 3: 143
Argynnis cornelia Edwards, 1892; Can. Ent. 24 (5): 106
Argynnis electa ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Argynnis cornelia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria atlantis electa ; [NACL, #4459o]
S. a. lurana dos Passos & Grey, 1945
Speyeria atlantis lurana dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Harney Peak, Black Hills, SD
Speyeria atlantis lurana ; [NACL, #4459p]
S. a. hutchinsi dos Passos & Grey, 1947
hutchinsi (Gunder, 1932); (ab.)
Speyeria atlantis hutchinsi ; [NACL, #4459q]
S. a. beani (Barnes & Benjamin, 1926)
Speyeria atlantis beani ; [NACL, #4459r]
S. a. lais (Edwards, 1883)
Alberta, British Columbia Argynnis lais Edwards, 1883; Can. Ent. 15 (11) : 209
Speyeria atlantis helena dos Passos & Grey, 1957; J. N.Y. Ent. Soc. 63: 95 (repl. Argynnis lais Edwards, 1883)
Argynnis lais ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 13
Speyeria lais ; [BOW: pl. 16, f. 3]
Speyeria atlantis lais ; [NACL, #4459s]
S. a. dennisi dos Passos & Grey, 1947
dennisi (Gunder, 1927); (ab.)
Speyeria atlantis dennisi ; [NACL, #4459t]
S. a. elko Austin, 1983
Nevada (Elko Co.) Speyeria atlantis elko Austin, 1983; J. Lep. Soc. 37 (3): 244, f. 1-2, TL: Nevada, Elko Co., Owybee River Valley, Wild Horse Creek Camping, ca. 10mi S of Mountain City

Speyeria hydaspe (Boisduval, 1869)
W.USA. See [About maps] Hydaspe Fritillary
Argynnis hydaspe Boisduval, 1869; Ann. Soc. ent. Belg. 12 : 60
Speyeria hydaspe, E.H. Strickland Ent. Mus, U of [ALBERTA]
Speyeria hydaspe, Butterflies of America [BOA]
Speyeria hydaspe, Butterflies and Moths of North America [BAMONA]
Speyeria hydaspe ; [NACL, #4460]; [EBW]; [Opler]
[hydaspe-1v.jpg] 800x828 (~114Kb) underside USA: Taneum Creek, Wenatchee National Forest, (47°06'24N 120°54'44W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela [hydaspe-2uU.jpg] identification uncertain 800x758 (~67Kb) upperside USA: FR9705, Table Mtn, Kittitas Co., WA, 30.7.1999, Photo © Markku Savela [hydaspe-3m.jpg] 500x673 (~59Kb) male USA: End of FR 6520 /Johnson Ridge, Mt. Baker Snoqualmie Nat. Forest (47°47'48N 121°15'45W), Snohomish Co., WA, 7.8.2001, Photo © Markku Savela
[hydaspe-3v.jpg] 500x582 (~46Kb) underside USA: End of FR 6520 /Johnson Ridge, Mt. Baker Snoqualmie Nat. Forest (47°47'48N 121°15'45W), Snohomish Co., WA, 7.8.2001, Photo © Markku Savela

Larva on Viola [NAB]


S. h. virdicornis (Comstock, 1925)
Speyeria hydaspe virdicornis ; [NACL, #4460a]
S. h. purpurascens (H. Edwards, 1877)
California, ... Argynnis monticola var. purpurascens H. Edwards, 1877; Proc. Calif. Acad. Sci. 7 : 170
caliginosa (Comstock, 1925)
Argynnis monticola purpurascens ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria hydaspe purpurascens ; [NACL, #4460b]
S. h. minor dos Passos & Grey, 1947
Speyeria (Speyeria) hydaspe minor dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 21
Argynnis hydaspe rhodope f. alt. minor McDunnough, 1927; Can. ent. 59 : 155, TL: Mt. McLean, B. C.
Speyeria hydaspe minor ; [NACL, #4460c]
S. h. rhodope (Edwards, 1874)
Vancouver, British Columbia hydaspe rhodope (Edwards, 1874); Trans. amer. ent. Soc. 5 (1): 13, TL: Cariboo District, British Columbia
gregsoni (Gunder, 1932)
Speyeria hydaspe rhodope, Butterflies of America [BOA]
Argynnis rhodope ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 14
Speyeria hydaspe rhodope ; [NACL, #4460d]
S. h. sakuntala (Skinner, 1911)
skinneri (Holland, 1931)
Speyeria hydaspe sakuntala ; [NACL, #4460e]
S. h. conquista dos Passos & Grey, 1945
Speyeria hydaspe conquista dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17), TL: Little Tesuque Canyon, nr Sante Fe, New Mexico, 8000ft
Speyeria hydaspe conquista ; [NACL, #4460f]

Speyeria mormonia (Boisduval, 1869)
California, Nevada. See [About maps] Mormon Fritillary
Argynnis mormonia Boisduval, 1869; Ann. Soc. ent. Belg. 12 : 58
Speyeria mormonia, E.H. Strickland Ent. Mus, U of [ALBERTA]
Speyeria mormonia mormonia, Butterflies of America [BOA]
Argynnis mormonia, NSG Voucher Specimen [Wahlberg]
Speyeria mormonia, Butterflies and Moths of North America [BAMONA]
Speyeria mormonia ; [NACL, #4461]; [EBW]; [BOW: pl. 16, f. 4]; [Opler]

Larva on (eggs near) Viola adunca? [NAB]

S. m. bischoffii (Edwards, 1870)
Alaska mormonia bischoffii (Edwards, 1870); Trans. amer. ent. Soc. 3 : 189, TL: Anchorage, Alaska
Argynnis eurynome bischoffi ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria mormonia bischoffii ; [NACL, #4461a]; Grey, 1989, J. Lep. Soc. 43 (1): 6
S. m. opis (Edwards, 1874)
British Columbia mormonia opis (Edwards, 1874); , TL: Bald Mt., Cariboo Dist., British Columbia
jesmondensis (McDunnough, 1940); (infrasubsp.)
= ; Grey, 1989, J. Lep. Soc. 43 (1): 6
Speyeria mormonia jesmondensis dos Passos & Grey, 1947
Speyeria mormonia opis ; [NACL, #4461b]
Speyeria mormonia jesmondensis ; [NACL, #4461c]
S. m. washingtonia (Barnes & McDunnough, 1913)
Argynnis bischoffi[sic] washingtonia Barnes & McDunnough, 1913; Contr. nat. Hist. Lep. N. Am. 2 (3): (91-164) , TL: Mt Ra[i]nier, 7000ft, [Pierce Co.], Wash[ington]
Argynnis eurynome washingtonia f. ranierensis Gunder, 1932; Can. Ent. 64 (2): (276-284), TL: Paradise Valley, Mount Ra[i]nier, Pierce Co., Washington
ranierensis ; Guppy & Kondla, 2002, J. Lep. Soc. 56 (3): 181
Speyeria rainierensis[sic] ; [NACL, #4461]
Speyeria mormonia washingtonia ; [NACL, #4461d]
S. m. erinna (Edwards, 1883)
Argynnis eurynome var. erinna Edwards, 1883; Can. Ent. 15 (2) : 33
cunninghami (Owen, 1893); Ent. News 4 : 246
Speyeria mormonia erinna, Butterflies of America [BOA]
Argynnis eurynome cunninghami ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Argynnis eurynome erinna ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria mormonia erinna ; [NACL, #4461e]

Larva on Viola palustris, V. adunca var. bellidifolia Hammond & Dornfeld, 1983, J. Lep. Soc. 37 (2): 120

S. m. arge (Strecker, 1878)
mormonia arge (Strecker, 1878); Synon. Cat. Macroplep.: 114
Argynnis astarte Edwards, 1862; (preocc. by Doubleday, 1847)
Argynnis montivaga Behr, 1864; (preocc. by Behr, 1863)
Argynnis eurynome arge ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria mormonia arge ; [NACL, #4461f]
S. m. artonis (Edwards, 1881)
Argynnis artonis Edwards, 1881; Trans. Amer. Ent. Soc. 9 : 1
rubyensis (Gunder, 1932)
Speyeria mormonia artonis, Butterflies of America [BOA]
Argynnis eurynome artonis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria mormonia artonis ; [NACL, #4461g]
S. m. eurynome (Edwards, 1872)
???, Montana, Alberta, Pacific Coast Argynnis eurynome Edwards, 1872; Trans. amer. ent. Soc. 4 : 66
Argynnis clio Edwards, 1874; Trans. amer. ent. Soc. 5 (2): 106
Argynnis eurynome ab. eris Igel, 1922; (preocc. by Meigen, 1828)
benjamini (Gunder, 1927)
Argynnis eurynome f. brucei Gunder, 1927
Argynnis eurynome clio f. gunderi Field, 1936; (preocc. by Comstock, 1925)
Speyeria mormonia eurynome ab. igeli dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: (1-30) (repl. Argnynnis eurynome ab. eris Igel, 1922)
Speyeria mormonia eurynome ab. fieldi dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: (1-30) (repl. Argynnis eurynome clio trans.f. gunderi Field, 1936)
Speyeria mormonia eurynome, Butterflies of America [BOA]
eurynomes ; Edwards, 1873, (missp.)
Argynnis eurynome ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Argynnis eurynome clio ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15
Speyeria mormonia eurynome ; [NACL, #4461h]
Speyeria eurynome ; [BOW: pl. 15, f. 18]
S. m. luski (Barnes & McDunnough, 1913)
Speyeria mormonia luski, Butterflies of America [BOA]
Speyeria mormonia luski ; [NACL, #4461i]

 


19.5.2013 (19)

References:
[ALBERTA]
University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU2]
Tuzov (ed.), Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000; Guide to the Butterflies of Russia and adjacent territories; Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae; Volume 2
[Bruce Walsh]
Bruce Walsh; e-mail: Bruce Walsh; The Moths of Southeastern Arizona; Butterflies of Southeastern Arizona
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[GALEPS]
Moths and Butterflies of Georgia and the Southeastern United States; James K. Adams; Dept. of Natural Science, Dalton State College, Dalton, GA ; http://www.daltonstate.edu/galeps/index.htm
[ILMB]
Interactive Listing of Mexican Butterflies
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Michael Borsch]
Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes
[Michel Tarrier]
Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org
[NAB]
Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[NORLEP]
Norges Sommerfugler
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[Pertti Pakkanen]
e-mail: Pertti Pakkanen; Butterfly and moth observations from Finland
[SPRK]
Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Tero Piirainen]
Tampere, Finland; e-mail: Tero Piirainen
[Vitaly Charny]
Birmingham, Alabama, USA
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Austin, 1983
A new subspecies of Speyeria atlantis from the great basin of Nevada J. Lep. Soc. 37 (3): 244-248
Barnes & McDunnough, 1913
New North American Lepidoptera with notes on described species Contr. nat. Hist. Lep. N. Am. 2 (3) : 93-165, pl. 1-9, I-IX
Behr, 1862
[Our California Argynnides] Proc. Cal. Acad. Sci. 2 : 172-177
Behr, 1863
On Californian Lepidoptera - No. III Proc. Calif. Acad. Sci. 3 (1) : 84-93
Boisduval, 1852
Lépidoptères de la Californie Ann. Soc. ent. Fr. (2) 10 : 275-324
Boisduval, 1869
Lépidoptères de la Californie Ann. Soc. ent. Belg. 12 : 5-28,37-94
Brown, 1983
The type of Argynnis apacheana Skinner J. Lep. Soc. 37 (1): 79-80
Chermock & Frechin, 1947
A new Speyeria from Washington Pan-Pacif. Ent. 23 (3): 111-112
Cockerell, 1900
A New Variety of Argynnis Nitocris Ent. News 11 (10) : 622
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
de Nicéville, 1889
On new and little-known Butterflies from the Indian Region, with Revision of the Genus Plesioneura of Felder and of Authors J. Bombay nat. Hist. Soc. 4 (3) : 163-194, pl. A-B
dos Passos, 1935
Some butterflies of southern Newfoundland with descriptions of new subspecies Can. Ent. 67: 82-88
dos Passos & Grey, 1942
Two new North American subspecies of Argynnis, with some revisional notes Amer. Mus. Novit. 1214: 6pp, 1fig
dos Passos & Grey, 1945
A new species and some new subspecies of Speyeria Amer. Mus. Novit. 1297: 17pp, 30fgs
dos Passos & Grey, 1947
Systematic catalogue of Speyeria with designations of types and fixations of type localities Amer. Mus. Novit. 1370: 30pp
dos Passos & Grey, 1957
A new name for Argynnis lais Edwards J. N.Y. Ent. Soc. 63: 95-96
Drury, [1773]
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 1 : 1-130, pl. 1-50 (1770), (1773)
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1862
Descriptions of certain Species of Diurnal Lepidoptera found within the limits of the United States and of British America. - No. [1]-3 Proc. Acad. nat. Sci. Philad. 13: 160-164 (1861), 14: 54-58, 221-226 (1862)
Edwards, 1864
Descriptions of certain species of Diurnal Lepidoptera, found within the limits of the United States and British America. (No. 1-3) Proc. ent. Soc. Philad. 2 (1): 14-22, pl. 1, 11 (1863), (2): 78-82 (1863), (4): 501-507 (1864)
Behr in Edwards, 1864; Edwards, 1864
Notes on the Argynnides of California Proc. ent. Soc. Philad. 3 : 434-436
Edwards, 1869
The Butteflies of North America. Vol. I. Butts. N. Amer. 1: 218pp, 50pls
Edwards, 1890
The Butteflies of North America. Vol. III. Butts N. Amer. 3: 431pp, 51pls
Edwards, 1869
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 2 : 369-376
Edwards, 1870
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 3 (1) : 10-22
Edwards, 1870
Descriptions of new North American diurnal Lepidoptera Trans. amer. ent. Soc. 3 : 189-196
Edwards, 1872
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 4 : 61-70
Edwards, 1874
Descriptions of new species of diurnal Lepidoptera found in North America Trans. amer. ent. Soc. 5 (1): 13-19, (2): 103-111
Edwards, 1876
Description of new species of diurnal Lepidoptera found within the United States and British North America Trans. amer. ent. Soc. 5 : 202-208
Edwards, 1876
New species of diurnal Lepidoptera Trans. amer. ent. Soc. 5 : 289-292
Edwards, 1878
Descriptions of new species of North American Lepidoptera Field & Forest 3: 115-119, 142-144
Edwards, 1879
Descriptions of new species of butterflies collected by Mr. H. K. Morrison in Nevada, 1878; also, Remarks on some errors of synonymy and arrangement Can. Ent. 11 (3): 49-56, : 79-80
Edwards, 1879
Descriptions of new species of North American butterflies; also, Notes upon certain species Can. Ent. 11 (5) : 81-89
Edwards, 1881
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. Amer. Ent. Soc. 9 : 1-8
Edwards, 1883
Descriptions of new species of diurnal Lepidoptera found in British America and the United States Can. Ent. 15 (2) : 32-36
Edwards, 1883
Descriptions of new species of North American butterflies Can. Ent. 15 (11) : 209-211
Edwards, 1886
Description of new species of butterflies found in the United States Can. Ent. 18 (4) : 61-65
Edwards, 1892
Miscellaneous notes on butterflies, larvae, etc. Can. Ent. 24 (3): 49-56, (5): 105-111
H. Edwards, 1877
Pacific Coast Lepidoptera. - No. 22. Notes on some Diurnal Lepidoptera, with description of new varieties Proc. Calif. Acad. Sci. 7 : 163-173
H. Edwards, 1877
Pacific Coast Lepidoptera. No. 26. - Description of two new forms of Diurnals Proc. Cal. Acad. Sc.: 1 (Dec. 17)
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus. Syst. Ent.: 832pp
Fruhstorfer, 1903
Neue Nymphaliden aus Ostasien Dt. ent. Z. Iris 16 (1) : 22-42, pl. 1, f. 5-6 (early October)
Fruhstorfer, 1908
Eine neue Argynnis-Rasse aus Spanien Ent. Zs. 22 (39) : 161 (26 December)
Godman & Salvin, [1901]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Grey, 1989
Sundry Argynnine concepts revisited J. Lep. Soc. 43 (1): 1-10
Gunder, 1924
Several New Aberrant Lepidoptera (Rhopalocera) from California Ent. News 35 (5) : 153-158
Gunder, 1925
Several New Varieties of and Aberrant Lepidoptera (Rhopalocera) from California Ent. News 36 : 1-9, pl. 1
Gunder, 1932
New Rhopalocera Can. Ent. 64 (2): 276-284
Guppy & Kondla, 2002
The correct spellings and type localities of butterflies named from Mount Rainier, Washington, USA J. Lep. Soc. 56 (3): 180-131
Hammond & Dornfeld, 1983
A new subspecies of Speyeria egleis from the pumice region of central Oregon J. Lep. Soc. 37 (2): 115-120
Holland, 1900
A Description of a Variety of Argynnis Nitocris from Chihuahua, Mexico Ent. News 11 (1) : 332-333
Kardakov, 1928
Zur Kenntnis der Lepidopteren des Ussuri-Gebietes Entomol. Mitt. 17 (4): 261-273
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Linnaeus, 1767
Systema Naturae per Regna tria Naturae, secundum Classes, Ordines, .... Editio Duocecima Reformata. Tom. 1. Part II. Syst. Nat. (Edn 12) 1 (2): -,533-1327
Matsumura, 1908
Die Nymphaliden Japans Ent. Zs. 22 (39) : 157-158
Matsumura, 1911
Erster Beitrag zur Insekten Fauna von Sachalin J. Coll. Agr. Tohoku Imp. Univ. Sapporo 4 (1) : 1-145, pl. 1-2
Matsumura, 1928
New Butterflies especially from the Kuriles Insecta Matsumurana 2 (4) : 191-201
Matsumura, 1929
Some new butterflies from Korea received from Mr. T. Takamuku Insecta Matsumurana 3 (4) : 152-156
McCorkle & Hammond, 1988
Biology of Speyeria zerene hippolyta in marine-modified environment J. Lep. Soc. 42 (3): 184-195
McDunnough, 1927
The Lepidoptera of the Seton Lake Region, British Columbia Can. ent. 59 : 152-162,193-199,207-214,239-246,266-277
McDunnough, 1935
A new race of Argynnis cybele from Nova Scotia Can. ent. 67: 18-19
Ménétriés, 1832
Catalogue raisonne des objects de zoologie recuelis dans un voyage au Caucase et jusqu'aux frontières de la Perse Cat. Rais.: -
Moore, [1875]
Descriptions of new Asiatic Lepidoptera Proc. zool. Soc. Lond. 1874 (4) : 565-579, pl. 66-67
Owen, 1893
Peculiar form of Argynnis erinna Edw Ent. News 4 : 246
Reakirt, 1866
Coloradian butteflies Proc. Ent. Soc. Philad. 6 (1) : 122-151
Reuss, 1926
Systemischer uberlich der Dryadinae T. Rss mit einigen Neubeschreibungen Dt. ent. Z. 1926 (1) : 65-70
Rothschild, 1933
On a collection of Lepidoptera from Spanish Morocco Novit. Zool. 38 : 315-330
Röber, 1896
Neue Schmetterlinge aus dem cilicischen Taurus Ent. Nachr. 22 (6) : 81-84
Scudder, 1872
A systematic Revision of some of the American Butterflies; with brief notes on those known to occur in Essex County, Mass. 4th Ann. Rep. Peabody Acad. Sci. (1871): 24-83
Scudder, 1875
Synonymic list of the butterflies of North Amercia, north of Mexico. (1) Nymphales Bull. Buffalo Soc. nat. Sci. 2 : 233-269
Skinner, 1897
Notes on Rhopalocera, with descriptions of new species and varieties Can. Ent. 29 (6) : 154-156
Skinner, 1917
New species of Lepidoptera Ent. News 28 (7) : 328-329
Skinner, 1918
Argynnis apacheana, a New Name (Lepid.) Ent. News 29 (2) : 67
Strecker, 1878
Butterflies and Moths of North America Synon. Cat. Macroplep.: -
Williams, 2002
Recognition of western populations of Speyeria idalia as new subspecies J. Lep. Soc. 55 (4): 144-149
Wolcott, 1916
Description of two Hitherto Undescribed Aberrations, the one of Argynnis alcestis, the of the Chlorippe celtis (Lep.) Ent. News, 27 (3) : 97-99, pl. 4


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.