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Coenonympha Hübner, [1819]

Coenonympha Hübner, [1819]; Verz. bek. Schmett. (5): 65, TS: Papilio geticus Esper
Chortobius [Dunning & Pickard], 1858; Accentuated List Brit. Lep.: 5, TS: Papilio pamphilus Linnaeus

= ; [NL4A, 223]

Chortobius Doubleday, 1859; Zoologist syn. List Brit. Butts (ed. 2): 2 (preocc. [Dunning & Pickard], [1859])

= ; [NL4A, 223]

Sicca Verity, 1953; Le Farfalle diurn. d'Italia 5: 83, TS: Papilio dorus Esper

= ; [NL4A, 223]

Coenonympha, NSG Voucher Specimen [Wahlberg]
Coenonympha (Coenonymphina) ; [Wahlberg]; [NL4A, 223]

Coenonympha pamphilus (Linnaeus, 1758)

Palearctic. See [About maps] Keltaniittyperhonen Small Heath Kamgräsfjäril Staggängefjäril Wiesenvögelchen Kleiner Heufalter Kälberauge Le Procris Nispola Strzępotek ruczajnik
Papilio pamphilus Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 472, TL: Sweden
?Coenonympha pamphiloides Reakirt, 1866; Proc. Ent. Soc. Philad. 6 (1) : 146

= ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30

?nephela Hufnagel

= ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30

?menalcas Poda, 1761; Ins. Mus. Graecensis: 78

= ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30

?Coenonympha pamphilus ab. semilyllus Krulikowsky, 1908; Soc. Ent. 23 (1): 3, TL: Wiatka; Kasan
Coenonympha pamphilus, NSG Voucher Specimen [Wahlberg]
Coenonympha pamphilus, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Coenonympha pamphilus, Svenska fjärilar [SVF]
Coenonympha pamphilus, Norges Sommerfugler [NORLEP]
Coenonympha pamphilus, Moths and Butterflies of Europe [MBE]
Coenonympha pamphilus, Europaische Schmetterlinge [Mario Maier]
Coenonympha pamphilus, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha pamphilius[sic] ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30
Coenonympha pamphilus ; [BRU, 192]; [EBW]; [BOW: pl. 6, f. 5]; [MRS, 404]; [Otakar Kudrna]

500x376 (~39Kb) underside Hoe lake, Devon, UK 24/06/95, Photo © Simon Coombes	1000x1076 (~169Kb) underside Russia, Tatarstan, Varkljed-Bodja (56°13'N 52°49'E), about 20km S of Agriz, 8.8.2004, Photo © Markku Savela	700x572 (~77Kb) underside France, Sophia Antipolis, 5.10.2000, Photo © Markku Savela
700x603 (~74Kb) underside France, Sophia Antipolis, 5.10.2000, Photo © Markku Savela	470x384 (~99Kb) underside Czech Republic, Blansko, 15.6.2004, Photo © Michal Koupý	1200x900 (~187Kb) underside Italy, Ravenna, August 2002, Photo © Maurizio Elia
293x244 (~10Kb) female	282x259 (~10Kb) male	900x723 (~54Kb) underside FINLAND, Ab: Suomusjärvi, 669:31, 28.6.1998 Photo © Markku Savela

Larva on Poaceae, Poa, Nardus, Stipa, Cynosurus, Dactilis, Festuca, Anthoxanthum, Brachypodium, (etc.) [BRU] Anthoxanthum odoratum, Deschampsia caespitosa, D. flexuosa [SPRK]

Various forms described, real status uncertain [BRU]

C. p. lyllus (Esper, 1806)

Papilio lyllus Esper, 1805; Die Schmett., Th I, Suppl. Th. 2 (11): 23, pl. 122, f. 1-2, TL: Portugal
Coenonympha pamphilus latevittata Verity, 1916; Ent. Rec. J. Var. 28 (7-8): 172, TL: Africa [Algeria]
Coenonympha pamphilus arenosa Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 198, TL: Fez (Morocco)
Coenonympha pamphilus atlantea Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 199, TL: Atlas Mts. (Morocco)
Coenonympha pamphilus lyllus ; [BMAT: 84, pl. 28, f. 28-44]

600x435 (~30Kb) underside Morocco, Pays Beni-Routen, Rif, Photo © Michel Tarrier

C. p. marginata Heyne, [1894]

pamphilus marginata Heyne, [1894]; in Rühl, Die palaearktischen Grossschmetterlinge und ihre Naturgeschichte 1 (Tagfalter): 619

C. p. fulvolactea Verity, 1926

pamphilus fulvolactea Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 205

C. p. centralasiae Verity, 1926

pamphilus centralasiae Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 205

C. p. infrarasa Verity, 1926

pamphilus infrarasa Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 206

C. p. juldusica Verity, 1926

pamphilus juldusica Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 206

C. p. ferghana Stauder, 1924

pamphilus ferghana Stauder, 1924; Int. ent. Z. 17: 152

C. p. nitidissima Verity, 1926

pamphilus nitidissima Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 199

C. p. asiaemontium Verity, 1926

pamphilus asiaemontium Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 206

Coenonympha saadi Kollar, [1849]

Iraq, Iran, Kurdistan. See [About maps] Coenonympha saadi Kollar, [1849]; Denkschr. Akad. Wiss. Wien. 1 : 52, TL: Shiraz, S.Iran
Coenonympha saadi, NSG Voucher Specimen [Wahlberg]
Coenonympha saadi ; [BOW: pl. 200, f. 2]

Larva on Poa, (and other grasses) [BRU, Hesselbarth, 1983]

C. s. iphias Eversmann, 1851

Transcaucasia saadi iphias Eversmann, 1851; Bull. Soc. imp. Nat. Moscou 24 (2) : 618
mesopotamica Heyne, [1894]; in Rühl, Die palaearktischen Grossschmetterlinge und ihre Naturgeschichte 1 (Tagfalter): 617

Coenonympha mangeri O. Bang-Haas, 1927

Afghanistan. See [About maps] mangeri O. Bang-Haas, 1927; Horae Macrolep. Palaearct. 1: 50

Coenonympha haydenii (Edwards, 1872)

Montana, Colorado. See [About maps] Hayden's Ringlet Wyoming Ringlet Yellowstone Ringlet
haydenii (Edwards, 1872); in Hayden, Geol. Surv. Montana: 467
Coenonympha haydenii, Butterflies and Moths of North America [BAMONA]
Coenonympha haydenii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30; [NACL, #4581]
Coenonympha haydeni ; [BOW: pl. 18, f. 14]; [Opler]

Larva on Poaceae?, Cyperaceae? [NAB]

All of these may just be single C. tullia.

Coenonympha tullia (Müller, 1764)

NWEU, Temperate Asia, NA. See [About maps] Saraikkoniittyperhonen Large Heath Common Ringlet Starrgräsfjäril Starrängefjäril Le Daphnis Grosser Heufalter Strzępotek soplaczek
Papilio tullia Müller, 1764; Fauna Ins. Fridr.: 36, TL: Zealand Is., Denmark
Papilio davus Fabricius, 1777; Genera Insectorum : 259
isis Thunberg, 1791; Diss. Ent. sistens Insecta Suecica (2): 31
Coenonympha tullia, E.H. Strickland Ent. Mus, U of [ALBERTA]
Coenonympha tullia, NSG Voucher Specimen [Wahlberg]
Coenonympha tullia, Butterflies and Moths of North America [BAMONA]
Coenonympha tullia, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Coenonympha tullia, Svenska fjärilar [SVF]
Coenonympha tullia, Norges Sommerfugler [NORLEP]
Coenonympha tullia, Moths and Butterflies of Europe [MBE]
Coenonympha tullia, Europaische Schmetterlinge [Mario Maier]
Coenonympha tullia ; [BRU, 193]; [EBW]; [BOW: pl. 6, f. 6]; [Opler]; [Otakar Kudrna]

599x400 (~31Kb) larva Central Europe 1990-2000, Photo © Michael Borsch	400x599 (~27Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch	400x588 (~38Kb) underside Central Europe 1990-2000, Photo © Michael Borsch
444x358 (~40Kb) underside USA: Alabama, 2.8.1999, Photo © Vitaly Charny	351x621 (~35Kb)	360x285 (~19Kb) female
370x282 (~21Kb) male

Larva on Eriophorum, Carex, Ryhchospora, Festuca, Poa, Stipa [BRU]

C. t. tullia (Müller, 1764)

EU, W.Siberia Coenonympha typhon Rottemburg, 1775; Der Naturforscher 6 : 15
?laidon (Borkhausen, 1788); Naturges. Eur. Schmett., 1: 91
timanensis Sedykh, 1977
?Coenonympha typhon ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30
?Coenonympha typhon laidon ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30

C. t. suecica Hemming, 1936

NEU, N.Siberia (Yamal Peninsula) fridolini Kusnezov, 1941; in Davenport
minimus Sedykh, 1977

C. t. bosniae Davenport, 1941

Coenonympha tullia bosniae Davenport, 1941; Bull. Mus. comp. Zool. Harv. 87 (4): 244-245, TL: Lake Jesero, Bosnia
bosniae ; Abadjiev, 2002, Neue Ent. Nachr. 23: 27

C. t. chatiparae Sheljuzhko, 1937

Caucasus Minor

C. t. elwesi Davenport, 1941

Altai Mts.

413x400 (~45Kb) underside male An alpine meadow, at 2200 m above sea level, on the eastern spurs of the Katunskiy Mt. Range, the valley of the Argem (Direntai) stream, Central Altai Mts., West Siberia, Russia. 13th July, 1988, Photo © Oleg Kosterin

C. t. sibirica Davenport, 1941

Amur Region

C. t. viluiensis Ménétriés, 1859

Chukot Peninsula witimensis Davenport, 1941

C. t. mixturata Alpheraky, 1897

Kamchatka Coenonympha kodiak mixturata ; [NACL, #4582a]

C. t. yontocket Porter & Mattoon, 1989

N.California (Del Norte Co.) Coenonympha tullia yontocket Porter & Mattoon, 1989; J. Lep. Soc. 43 (3): 231, f. 1-3, TL: California, Del Norte Co. (dunes N of Crescent City, between Lake Earl and Smith River)

Coenonympha kodiak Edwards, 1869

Alaska, Yukon, NW.Territories, N.Siberia. See [About maps] Kodiak Ringlet
Coenonympha kodiak Edwards, 1869; Trans. amer. ent. Soc. 2 : 375
Coenonympha kodiak ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30; [NACL, #4582]

Larva on Poaceae?

C. k. yukonensis Holland, 1900

Coenonympha kodiak var. yukonensis Holland, 1900; Ent. News 11 (3): 386
Coenonympha kodiak yukonensis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30; [NACL, #4582b]

Coenonympha inornata Edwards, 1861

Newfoundland, Canada, Montana. See [About maps] Prairie Ringlet
quebecensis Barnes & Benjamin, 1926
Coenonympha tullia inornata ; [USGS]
Coenonympha inornata ; [NACL, #4583]; [BOW: pl. 18, f. 7]

Larva on Poaceae [NAB]

C. i. mcisaaci dos Passos, 1935

Coenonympha inornata mcisaaci dos Passos, 1935; Can. Ent. 67: (82-88), TL: Doyles Station, Newfoundland
maciasaaci dos Passos, 1964; (emend.)
Coenonympha inornata mcisaaci ; [NACL, #4583a]

C. i. benjamini McDunnough, 1928

Coenonympha inornata benjamini ; [NACL, #4583b]

C. i. nipisquit McDunnough, 1939

New Brunswick, Quebec Nipisquit Ringlet
Coenonympha inornata[?] nipisquit McDunnough, 1939; Can. ent. 71: 266, TL: near Bathurst, New Brunswick
Coenonympha inornata nipisquit ; [NACL, #4583c]
Coenonympha tullia nipisquit ; Webster, 1998, J. Lep. Soc. 52 (4): 345-355
Coenonympha nipisquit ;
?nipisiquit ; (missp.?)

Larva on Spartina patens, less Festuca rubra Webster, 1998, J. Lep. Soc. 52 (4): 350

C. i. heinemanni F.M. Brown, 1959

Coenonympha inornata heinemanni ; [NACL, #4583d]

Coenonympha ochracea Edwards, 1861

Colorado. See [About maps] Ocher Ringlet
Coenonympha ochracea Edwards, 1861; Proc. Acad. nat. Sci. Philad. 13: 161, TL: Colorado
Coenonympha albescens Field, 1936
Coenonympha phantasma Burdick, 1956
Coenonympha ochracea ; [NL4A, #1602]
ochracia F. & R. Chermock, 1938; (missp.)
Coenonympha ochracea ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30; [NACL, #4584]
Coenonympha tullia ochracea ; [USGS]

Larva on Poaceae? [NAB]

C. o. brenda Edwards, 1869

California Caenonympha[sic] ochracea brenda Edwards, 1869; , TL: California
Coenonympha ochracea brenda ; [NACL, #4584a]; [NL4A, #1602a]

C. o. mackenziei Davenport, 1936

Coenonympha ochracea mackenziei ; [NACL, #4584b]

C. o. mono Burdick, 1942

Coenonympha ochracea mono ; [NACL, #4584c]

C. o. furcae Barnes & Benjamin, 1926

Coenonympha ochracea furcae ; [NACL, #4584d]

C. o. subfusca Barnes & Benjamin, 1926

Coenonympha ochracea subfusca ; [NACL, #4584e]

Coenonympha ampelos Edwards, 1871

-. See [About maps] Northwest Ringlet
Coenonympha tullia ampelos ; [USGS]
Coenonympha ampelos ; [NACL, #4585]

identification uncertain 700x510 (~33Kb) underside USA: Lake Sammamish Park, Issaquah (47°33'36N 122°03'25W), King Co., WA, 11.7.1999, Photo © Markku Savela	identification uncertain 800x739 (~47Kb) underside USA: Issaquah (47°32'35N 122°04'00W), King Co., WA, 7.7.1999, Photo © Markku Savela	identification uncertain 500x439 (~34Kb) underside USA: FR9708, Table Mtn, Kittitas Co., WA, 19.7.1999, Photo © Markku Savela
identification uncertain 900x1002 (~120Kb) underside USA: near Tyler (47°27'09"N 117°48'41"W), Spokane Co., Washington, 18.8.2005, Photo © Markku Savela

Larva on Poaceae [NAB]

C. a. eunomia Dornfield, 1967

eunomia Field, 1937; (spring form.)
Coenonympha ampelos eunomia ; [NACL, #4585a]

C. a. elko Edwards, 1881

Nevada - Vancouver Coenonympha elko Edwards, 1881; Can. Ent. 13 (3) : 57
Coenonympa sweadneri F. & R. Chermock, 1941
Coenonympha elko ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30
Coenonympha ampelos elko ; [NACL, #4585b]
Coenonympha elko ; [BOW: pl. 18, f. 6]

C. a. columbiana McDunnough, 1928

Coenonympha ampelos columbiana ; [NACL, #4585c]

C. a. insulana McDunnough, 1928

Coenonympha ampelos insulana ; [NACL, #4585d]

Coenonympha california Westwood, [1851]

WNA. See [About maps] California Ringlet
Coenonympha california Westwood, [1851]; Gen. diurn. Lep. (2): 398, TL: California
californica H. Edwards, 1881; (missp.)
Coenonympha californica ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 29
Coenonympha california ; [NACL, #4586]; [EBW]; [BOW: pl. 18, f. 5]
Coenonympha tullia californica ; [USGS]
Coenonympha california ; [NL4A, #1601]

Larva on Poaceae [NAB]

C. c. california

California Satyrus galactinus Boisduval, 1852; , TL: California

= ; [NL4A, #1601a]

Coenonympha ceres Butler, 1866; , TL: California

= ; [NL4A, #1601a]

Coenonympha californica[sic] var. pulla H. Edwards, 1881; Papilio 1 (4) : 51, TL: San Mateo Co., California

= ; [NL4A, #1601a]

Coenonympha californica pulla ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 29
Coenonympha california california ; [NL4A, #1601a]

C. c. eryngii H. Edwards, 1877

siskiyouensis Comstock, 1925
Coenonympha california eryngii ; [NACL, #4586a]

Coenonympha caeca Staudinger, 1886

Pamirs-Alai, Ghissar, Tian-Shan - Altai. See [About maps] Coenonympha caeca Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 251, TL: Chatkalsky Mts, Uzbekistan
Coenonympha caeca ; [BRU, 193]

Larva on Carex spp. [BRU]

?C. c. eupompus (Stauder, 1924)

Dzhungarsky Alatau caeca eupompus (Stauder, 1924); , TL: Ili
pamphilus eupompus Stauder, 1924
Coenonympha caeca heptopotamica Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 352, TL: Aulie-ata

C. c. subcaeca Heyne, [1894]

Sailyughem Mts

Coenonympha symphita Lederer, 1870

NE.Turkey, SW.Georgia. See [About maps] Coenonympha symphita Lederer, 1870; Ann. Soc. ent. Belg. 13 : 44, TL: Between Akhaltsikhe and the Turkish border, Georgia
Coenonympha symphita karsiana Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 351, TL: Transcaucasia
ab. inocellata Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 352
Coenonympha symphita ; [BRU, 194]

Larva on Poa annua [BRU, Hesselbarth et al, 1995]

Coenonympha rhodopensis Elwes, 1900

Italy, Romania, Bulgaria, Albania, N.Greece, Yugoslavia. See [About maps] Eastern Large Heath
Coenonympha tiphon var. rhodopensis Elwes, 1900; Trans. ent. Soc. Lond. 1900 (2) : 205, TL: Bulgaria, Rila Mts., Levi Iskar, 5000ft
Coenonympha rhodopensis, NSG Voucher Specimen [Wahlberg]
Coenonympha rhodopensis, Moths and Butterflies of Europe [MBE]
Coenonympha rhodopensis, Captain's European Butterfly Guide [Simon Coombes]
rhodopensis ; Abadjiev, 2002, Neue Ent. Nachr. 23: 27
Coenonympha rhodopensis ; Abadjiev, 2000, Atalanta 31 (3/4): 461-466; [Otakar Kudrna]

Coenonympha corinna (Hübner, 1804)

Corsica, Sardinia, Elba. See [About maps] Corsican Heath
corinna (Hübner, 1804); Samml. eur. Schmett. [1] : f. 536, 537
Coenonympha corinna, NSG Voucher Specimen [Wahlberg]
Coenonympha corinna, Moths and Butterflies of Europe [MBE]
Coenonympha corinna ; [BOW: pl. 5, f. 16]; [Otakar Kudrna]

Coenonympha fettigii Oberthür, 1874

Morocco, Algeria, Tunisia. See [About maps] Coenonympha fettigii Oberthür, 1874; Petites Nouv. Ent. 1: 412, TL: Province of Oran [Telaghre] (Algeria)
Coenonympha fettigii, Moths and Butterflies of Europe [MBE]
Coenonympha dorus fettigii ;

C. f. fettigii

Coenonympha fettigii fettigii ab. infra-simplex Rothschild, 1917; Novit. Zool. 24 (1): 118, TL: Les Pins (Algeria)
Coenonympha fettigii fettigii ; [BMAT: 84, pl. 29, f. 9-11]

C. f. nicholasi Rothschild, 1925

Algeria (Blida, Chrea, etc.), Tunisia (Aïn Draham, Tunis) Coenonympha dorus nicholasi Rothschild, 1925; Novit. Zool. 32 : 208, TL: Blida-les-Glaciéres (Algeria)
Coenonympha fettigii holli Oberthür, 1910; Ètud. Lépid. Comp. 4 : 42, pl. 48, f. 396-397, TL: Blida-les-Glaciéres (Algeria)
Coenonympha fettigii nicholasi ; [BMAT: 85, pl. 29, f. 12-17]

C. f. inframaculata Oberthür, 1922

W.Morocco (Rif, Midle Atlas, High Atlas) Coenonympha fettigi var. inframaculata Oberthür, 1922; Étud. Lépid. Comp. 19 (1): 87, TL: Fort Toumliline (Morocco)
Coenonympha fettigii inframaculata ; [BMAT: 86, pl. 29, f. 18-31]

600x435 (~33Kb) underside Morocco, Djebel Tisiréne, Rif, Photo © Michel Tarrier

Coenonympha dorus (Esper, 1782)

NAF, SWEU, it. See [About maps] Dusky Heath Velago de negro
Papilio dorus Esper, 1782; Die Schmett. Th. I, Bd. 2 (5): 130, pl. 78 f. 1
Coenonympha mathewi Tutt, 1904; Ent. Rec. J. Var. 16 : 308, TL: Vigo
Coenonympha dorus, NSG Voucher Specimen [Wahlberg]
Coenonympha dorus, Moths and Butterflies of Europe [MBE]
Coenonympha dorus, Europaische Schmetterlinge [Mario Maier]
Coenonympha dorus, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha dorus ; [EBW]; [BOW: pl. 5, f. 17]; [Otakar Kudrna]

568x740 (~90Kb) Albarracin, prov. Teruel, , Spain, August-1988, Photo © Enrique Garcia-Barros

Larva on Agrostis spp., Festuca ovina [BMAT]

C. d. austauti Oberthür, 1881

W.Algeria Austaut's Algerian Heath
Coenonympha dorus var. austauti Oberthür, 1881; Étud. d'Ent. 6: 59, TL: Nemours [Ghazaouet] (Algeria)
Coenonympha dorus austauti ; [BMAT: 86, pl. 29, f. 1-8]

Coenonympha vaucheri Blachier, 1905

Morocco. See [About maps] Vaucher's Heath
Coenonympha vaucheri Blachier, 1905; Bull. Soc. ent. Fr. 1905 : 213, TL: High Atlas Mts. (Morocco)
Coenonympha vaucheri, NSG Voucher Specimen [Wahlberg]
Coenonympha vaucheri, Moths and Butterflies of Europe [MBE]
Coenonympha vaucheri, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha vaucheri ; [BOW: pl. 115, f. 6]

C. v. vaucheri

Morocco (High Atlas) Coenonympha vaucheri ab. geminipuncta Blachier, 1905; Bull. Soc. ent. Fr. 1905 : 213, TL: High Atlas Mts. (Morocco)
Coenonympha meade-waldoi Elwes, 1905; Proc. zool. Soc. Lond. 1905 (4): 377
Coenonympha vaucheri ab. mediocellis Le Cerf, 1923; Bull. Soc. ent. Fr. 1923: 224, TL: Dj.Tachdirt (High Atlas Mts.) (Morocco)
Coenonympha vaucheri vaucheri ; [BMAT: 86, pl. 29, f. 32-40]

599x434 (~51Kb) underside Morocco, Djebel Azourki, Haut Atlas, Photo © Jean Delacre

C. v. annoceuri Wyatt, 1952

Morocco (Middle Atlas) Coenonympha vaucheri annoceuri Wyatt, 1952; Z. wien. ent. Ges. 37: 175, pl. 21, f. B1-B4, TL: Annoceur (Morocco)
Coenonympha vaucheri annoceuri ; [BMAT: 87, pl. 29, f. 41-48]

C. v. rifensis Weiss, 1979

Morocco (W.Rif Mts.) Coenonympha vaucheri rifensis Weiss, 1979; Entomops 48: 271, f. 1, TL: Dj. Lakraa (Morocco)
Coenonympha vaucheri rifensis ; [BMAT: 87, pl. 29, f. 49-57]

C. v. beraberensis Lay & Rose, 1979

Morocco (Tizi-n'Ouguerd-Zegzaoune, Dj.Aourach, Imlil, Lake Tislit, etc.) Coenonympha vaucheri beraberensis Lay & Rose, 1979; Ent. Z. Frankf. a. M. 89 (13): 143, f. 2, TL: [Tizi-n'Ouguerd-Zegzaoune] (Morocco)
Coenonympha vaucheri beraberensis ; [BMAT: 87, pl. 29, f. 58-66]

600x435 (~34Kb) underside Morocco, Tizi-n-Inezgane (Imilchil), Haut Atlas, Photo © Michel Tarrier

Coenonympha arcania (Linnaeus, 1761)

EU, Asia Minor, S.Russia, S.Urals. See [About maps] Pearly Heath Pärlgräsfjäril Perlgrasfalter Mancha Leonada Strzępotek perełkowiec
Papilio arcania Linnaeus, 1761; Fauna Suecica (Edn 2): 273, TL: Sweden
Papilio amyntas Poda, 1761; Ins. Mus. Graecensis: 79
caucasica Jachontov, 1914²
Coenonympha arcania, NSG Voucher Specimen [Wahlberg]
Coenonympha arcania, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Coenonympha arcania, Svenska fjärilar [SVF]
Coenonympha arcania, Norges Sommerfugler [NORLEP]
Coenonympha arcania, Moths and Butterflies of Europe [MBE]
Coenonympha arcania, Europaische Schmetterlinge [Mario Maier]
Coenonympha arcania, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha arcania ; [BRU, 195]; [EBW]; [BOW: pl. 5, f. 15]; [MRS, 405]; [Otakar Kudrna]

500x471 (~45Kb) underside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes	652x452 (~27Kb) underside male Poland, Tresta near Tomaszow Mazowiecki, 07.1997, Photo © Marek Michalski leg.	309x300 (~24Kb) underside An opening in an oak forest, the town Krymsk environs, Krymsk District, Krasnodarskii Krai Province, S Russia. 26th May 1990, Photo © Oleg Kosterin
517x689 (~57Kb) underside Sweden, Ekebo, Julita, 5.7.2003, Photo © Leif Wahlberg	479x639 (~66Kb) underside Sweden, Ekebo, Julita, 20.6.2004, Photo © Leif Wahlberg	470x441 (~93Kb) underside Czech Republic, Blansko, 15.6.2004, Photo © Michal Koupý

Larva on Gramineae, Melica [H&R]

Coenonympha gardetta (de Prunner, 1798)

Alps, N.Balkans. See [About maps] Alpine Heath
gardetta (de Prunner, 1798); Lepid. Pedemont.: 73

= Coenonympha pamphilus ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30

Papilio satyrion Esper, 1805; Die Schmett., Th I, Suppl. Th. 2 (11): 24, pl. 122, f. 2
philea (Hübner, 1800)
neoclides (Hübner, 1805)
Coenonympha gardetta, NSG Voucher Specimen [Wahlberg]
Coenonympha gardetta, Moths and Butterflies of Europe [MBE]
Coenonympha gardetta, Europaische Schmetterlinge [Mario Maier]
Coenonympha gardetta, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha gardetta ; [Otakar Kudrna]

Coenonympha darwiniana Staudinger, 1871

SEU. See [About maps] Coenonympha darwiniana, NSG Voucher Specimen [Wahlberg]
Coenonympha darwiniana, Captain's European Butterfly Guide [Simon Coombes]

Coenonympha arcanioides (Pierret, 1837)

Morocco, Algeria, Tunisia. See [About maps] Moroccan Pearly Heath
Satyrus arcanioides Pierret, 1837; Ann. Soc. ent. Fr. 6 : 306, TL: Province of Oran (Algeria)
?Satyrus marginalis Butler, 1868; Cat. diurn. Lep. Satyridae Brit. Mus. : 42

= ?Coenonympha arcania ;

Coenonympha arcanioides f. major Seitz, 1908; Gross-Schmett. Erde 1 : 143, pl. 48[e], TL: Algeria, Morocco
Coenonympha arcanioides gen. aest. holli Oberthür, 1910; Ètud. Lépid. Comp. 4 : 20, TL: Algeria
Coenonympha arcanioides ab. biocellata Oberthür, 1910; Ètud. Lépid. Comp. 4 : 21, TL: [Algeria]
Coenonympha arcanioides f. pluriocellata Stetter-Stättermayer, 1933; Int. ent. Z. 27: 340, TL: Bône [Annaba] (Algeria)
Coenonympha arcanioides f. inocellata Stetter-Stättermayer, 1933; Int. ent. Z. 27: 340, TL: Bône [Annaba] (Algeria)
Coenonympha arcanioides f. obliterata Stetter-Stättermayer, 1933; Int. ent. Z. 27: 340, TL: Bône [Annaba] (Algeria)
Coenonympha arcanioides f. nigro-ocellata Stetter-Stättermayer, 1933; Int. ent. Z. 27: 340, TL: Bône [Annaba] (Algeria)
Coenonympha arcanioides, NSG Voucher Specimen [Wahlberg]
Coenonympha arcanioides, Moths and Butterflies of Europe [MBE]
Coenonympha arcanioides ; [BOW: pl. 5, f. 12]; [BMAT: 88, pl. 28, f. 45-49]

600x435 (~30Kb) underside Morocco, Chefchaouen, Rif, Photo © Michel Tarrier

599x434 (~19Kb) underside Morocco, Chefchaouen, Rif, Photo © Jean Delacre

Coenonympha leander (Esper, 1784)

Hungary, Bulgaria, S.Russia, Asia Minor, Armenia, Iran. See [About maps] Russian Heath
Papilio leander Esper, 1784; Die Schmett. Th. I, Bd. 2 (9): 176, pl. 89, f. 5, TL: Volga region, Russia
Coenonympha leander, NSG Voucher Specimen [Wahlberg]
Coenonympha leander, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha leander ; [BRU, 195]; [BOW: pl. 6, f. 3]; [Otakar Kudrna]

Larva on Poa, Melica, Anthoxanthum, Lolium [BRU, Koenig, 1959, Hesselbarth et al, 1995, etc.]

C. l. obscura Heyne, 1894

Caucasus Minor, Armenian Highland, Talysh

Coenonympha hero (Linnaeus, 1761)

CEU, NEU, Temperate Asia, Amur, Korea, Japan. See [About maps] Ruskoniittyperhonen Scarce Heath Wald-Wiesenvögelchen Brun gräsfjäril Madängefjäril Strzępotek hero
Papilio hero Linnaeus, 1761; Fauna Suecica (Edn 2): 274, TL: Sweden
Coenonympha hero, NSG Voucher Specimen [Wahlberg]
Coenonympha hero, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Coenonympha hero, Svenska fjärilar [SVF]
Coenonympha hero, Norges Sommerfugler [NORLEP]
Coenonympha hero, Moths and Butterflies of Europe [MBE]
Coenonympha hero, Europaische Schmetterlinge [Mario Maier]
Coenonympha hero ; [BRU, 196]; [EBW]; [BOW: pl. 6, f. 1]; [MRS, 404]; [Otakar Kudrna]

Larva on Gramineae, Elymus arenarius [H&R], Elymus, Poa, Hordeum, Calamagrostis [BRU, Lang, 1884, Henriksen, Kreutzer, 1982]

C. h. sabaeus Fabricius, 1775

E.Europe, Urals, W.Siberia

C. h. perseis Lederer, 1853

up to Pacific sibirica Fuchs, 1899
Coenonympha hero perseis ; [MRS, 404]

478x640 (~92Kb) upperside underside male Russia, N Tuva, Piykhemskiy distr., h=800., 06.1997 Vashchenko S.N. leg, Photo © D. Smirnov

Larva on Cyperaceae [MRS]

C. h. pilwonis Matsumura, 1925

Sakhalin

C. h. latefasciata Matsumura, 1925

Kunashir

C. h. coreana Matsumura, 1927

Korea Coenonympha hero coreana Matsumura, 1927; Insecta Matsumurana 1 (4) : 164, TL: Corea

Coenonympha glycerion (Borkhausen, 1788)

EU, S.Siberia, Japan. See [About maps] Idänniittyperhonen Chestnut Heath Rostbraunes Wiesvögelchen Darrgräsfjäril Darrängefjäril Strzępotek glicerion
Papilio glycerion Borkhausen, 1788; Naturges. Eur. Schmett., 1: 90
Papilio iphis Denis & Shiffermüller, 1775; (preocc.)
Coenonympha iphis var. anaxagoras Assmus, 1857; Breslau Ent. Z. 10 : 9
Papilio tiphon Esper, 1777; Die Schmett. Th. I, Bd. 1 (9): 341, (6): pl. 35, f. 3-4 (preocc.)
Coenonympha glycerion glycerion, Lepidoptera collection [SZMN]
Coenonympha glycerion, NSG Voucher Specimen [Wahlberg]
Coenonympha glycerion, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Coenonympha glycerion, Moths and Butterflies of Europe [MBE]
Coenonympha glycerion, Europaische Schmetterlinge [Mario Maier]
Coenonympha glycerion, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha glycerion ; [BOW: pl. 5, f. 18]; [MRS, 405]; [Otakar Kudrna]

500x430 (~28Kb) underside Finland, Ylöjärvi, Teivaala, 6828:320, 16.7.1996, Photo © Tero Piirainen	800x540 (~51Kb) underside Finland, Vihti, Nummela, ?July 2002, Photo 10072002_24 © Timo Viitanen	470x443 (~80Kb) underside Czech Republic, Blansko, 15.6.2004, Photo © Michal Koupý
470x379 (~76Kb) underside Czech Republic, Blansko, 22.7.2004, Photo © Michal Koupý	600x400 (~60Kb) underside Estonia, Hiiumaa, Kootsaare nina. 24.07.2004, Photo © Hannu Tanner	724x724 (~116Kb) underside Russia, Moscow area, 10.06.2007, Photo © D. Smirnov
710x522 (~96Kb) underside male Russia: Moscow area (36°25'E, 56°00'N), 14.7.2008, Photo © D. Smirnov	811x660 (~113Kb) underside female Russia, Moscow area, 18.8.2010 (36°25'E, 56°00'N), Photo © D. Smirnov	333x278 (~15Kb) female
320x275 (~23Kb) male

Larva on Gramineae, Deschampsia caespitosa [SPRK]

Larva on Briza, Poa, Brachypodium, Melica, Cynosurus, (and other grasses) [BRU, Eckstein, 1913, Nekrutenko, 1990, Gorbuno, Korshunov, 1995]

C. g. alta Sheljuzhko, 1937

Caucasus

C. g. korshunovi Nekrutenko, 1978

Crimea

C. g. iphicles Staudinger, 1892

N.Alai, Sayan, Transbaikalia, Amur, Ussuri heroides Christoph, 1893
Coenonympha glycerion iphicles, Lepidoptera collection [SZMN]

C. g. iphina Staudinger, 1892

Saur, Tarbagatai

C. g. wutaica Murayama

Coenonympha glycerion wutaica ; [MRS, 405]

Coenonympha mahometana Alphéraky, 1881

Tian-Shan. See [About maps] Coenonympha mahometana Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 428, TL: Kuldzha, Kunges Valley, W. China
Coenonympha mahometana, NSG Voucher Specimen [Wahlberg]
Coenonympha mahometana ; [BRU, 197]

Coenonympha sunbecca Eversmann, 1843

Ghissar - Tarbagatai. See [About maps] Coenonympha sunbecca Eversmann, 1843; Bull. Soc. Imp. Nat. Moscou 16 (3) : 538, pl. 7, f. 4a-b, TL: Zaisan, SE. Kazakhstan
?sunbecca var. alexandra Heyne, [1894]; , TL: Kirghizsky Mts.
Coenonympha sunbecca, NSG Voucher Specimen [Wahlberg]
Coenonympha sunbecca alexandra ;
Coenonympha sunbecca ; [BRU, 197]; [EBW]; [BOW: pl. 200, f. 7]; [MRS, 404]

300x400 (~28Kb) underside A steppefied meadow in the middle flow of the Kora River, 30 km NE of the Tekeli City (1900 m above sea level), the North Dzhungarskiy Alatau mountain chain (NE part of Tien Shan), Taldy-Kurgan Province, SE Kazakhstan. 15th June, 1993 (on Artemisia santolinifolia Turcz. ex Besser), Photo © Oleg Kosterin

Coenonympha iphioides Staudinger, 1870

Spain. See [About maps] Spanish Heath Castaneo morena
Coenonympha iphioides Staudinger, 1870; Berl. Ent. Z. 14 : 101
Coenonympha iphioides, NSG Voucher Specimen [Wahlberg]
Coenonympha iphioides, Moths and Butterflies of Europe [MBE]
Coenonympha iphioides ; [BOW: pl. 6, f. 2]; [Otakar Kudrna]

612x520 (~45Kb) underside Barbatona (prov. Guadalajara), Spain 1-august-1989, Photo © Enrique Garcia-Barros

Coenonympha oedippus (Fabricius, 1787)

CEU, S.Siberia - Ussuri, China, Korea, Japan. See [About maps] False Ringlet Moor-Wiesenvögelchen Lindos ojos Strzępotek edypus
Papilio oedippus Fabricius, 1787; Mantissa Insectorum 2: 31, TL: S.Russia
Papilio geticus Esper, 1794; Die Schmett., Suppl. Th 1 (3-4): 51, (5-6): pl. 102, f. 2
Coenonympha annulifer Butler, 1877; Ann. Mag. nat. Hist. (4) 19 (109) : 91, TL: 370 miles from Tokei (Yedo)

= ; [NHM]

Coenonympha oedippus; [BRU, 196]
Coenonympha oedippus, NSG Voucher Specimen [Wahlberg]
Coenonympha oedippus, Moths and Butterflies of Europe [MBE]
Coenonympha oedippus, Europaische Schmetterlinge [Mario Maier]
Coenonympha oedippus ; [BOW: pl. 6, f. 4]; [MRS, 403]; [Otakar Kudrna]

1200x1565 (~150Kb) underside Hungary: Ocsa, June 2005, Photo © Szabolcs Safian

Larva on Lolium, Carex, Iris pseudacorus [H&R], Poa [BRU], Phragmites communis [MRS]

C. o. magna Heyne, 1895

Amur, Ussuri oedippus amurensis Heyne, 1895; in Ruhl
Coenonympha oedippus amurensis Rühl; [MRS, 403]

600x800 (~97Kb) underside male female A meadow steppe on the hill Malyi Batur on the left bank of the Onon River 7 km upstream of the village Nizhnii Tsasuchei, Onon District, Chita Province, SE Transbaikalia (Dahuria), Siberia, Russia. 11th July 1996, Photo © Oleg Kosterin

C. o. taibaica Maurayama

Coenonympha oedippus taibaica ; [MRS, 403]

Coenonympha nolckeni Erschoff, 1874

Ghissar, Alai, Tian-Shan, N.Pamirs, W.Pamirs. See [About maps] Coenonympha nolckeni Erschoff, 1874; in Fedschenko, Travels in Turkestan. 2 (5) : 23, pl. 2, f. 17, TL: Mt. Naubid, Zeravshan Valley, Uzbekistan
Coenonympha nolckeni, NSG Voucher Specimen [Wahlberg]
Coenonympha nolckeni ; [BRU, 196]; [BOW: pl. 200, f. 9]

Coenonympha amaryllis (Stoll, [1782])

S.Urals, Altai Mts, W.Siberia, E.Siberia, Transbaikalia, Amur, Ussuri, China, Mongolia, Korea. See [About maps] Papilio amaryllis Stoll, [1782]; in Cramer, Uitl. Kapellen 4 (32-32): 210, pl. 391, f. A, B, TL: "...de la Sibérie"
rinda Ménétriés, 1859; (infrasubsp.) - Amur
accrescens Staudinger, 1901; (infrasubsp.) - Ussuri
borisovi Korshunov & Ivonin, 1996; (infrasubsp.) - Baikal region
Coenonympha amaryllis, Lepidoptera collection [SZMN]
Coenonympha amaryllis, NSG Voucher Specimen [Wahlberg]
Coenonympha amaryllis ; [BRU, 194]; [BOW: pl. 200, f. 5]; [MRS, 403]; [Otakar Kudrna]

400x398 (~41Kb) underside The valley of the Shivilig-Khem River just upstream its leaving the the southern foot of the East Tannu-Ola mounatin range for the Ubsu-Nur Hollow, Tes-Khem District, Tuva Republic, Siberia, Russia. 11th July 1990, Photo © Oleg Kosterin

600x800 (~94Kb) upperside A steppefied meadow on the Onon River right bank terrace at a pine forest edge, 5 km upstream of the village Nizhnii Tsasuchei, Onon District, Chita Province, Transbaikalia, Siberia, Russia, 30th June 1996 (Two individuals catched by a Thomisidae spider), Photo © Oleg Kosterin

Larva on Poa [BRU]

C. a. simingica Murayama

Coenonympha amaryllis simingica ; [MRS, 403]

Coenonympha xinjiangensis Chou & Huang, 1994

Xinjiang. See [About maps] Coenonympha xinjiangensis Chou & Huang, 1994; in Chou, Monographia Rhopalocerum Sinensium 1-2: 759, 403, f. 32-33, TL: Xinjiang, Jiandengyu

Coenonympha tydeus Leech, [1892]

Tibet. See [About maps] Coenonympha tydeus Leech, [1892]; Butts China Japan Corea (1): 96
Coenonympha tydeus ; [MRS, 405]

Coenonympha sinica Alphéraky, 1888

Kashmir, Tibet. See [About maps] Coenonympha sinica Alphéraky, 1888; Stettin Ent. Ztg 49 : 66, TL: Nian-Chian-Kette, Djin-ta-sy
Coenonympha sinica ; [MRS, 405]

Coenonympha pavonina Alphéraky, 1888

China. See [About maps] Coenonympha pavonina Alphéraky, 1888; Stettin Ent. Ztg 49 : 66, TL: R. Hei-ho

Coenonympha semenovi Alphéraky, 1887

Tibet, W.China. See [About maps] Coenonympha semenovi Alphéraky, 1887; in Romanoff, Mem. lép. 3 : 405
Coenonympha semenovi, NSG Voucher Specimen [Wahlberg]
?Coenonympha semenori[sic?] ; [BOW: pl. 200, f. 6]
Coenonympha semenovi ; [MRS, 404]

1000x1058 (~190Kb) underside female China, Qinghai prov, 20 km S Tianjun (3900m), Photo © A. Timchenko leg.

1000x915 (~154Kb) upperside male China, Qinghai prov, 20 km S Tianjun (3900m), Photo © A. Timchenko leg.

1000x1062 (~176Kb) underside male China, Qinghai prov, 20 km S Tianjun (3900m), Photo © A. Timchenko leg.

C. s. jiadengyuica Huang & Murayama, 1992

Xianjing Coenonympha semenovi jiadengyuica Huang & Murayama, 1992; Tyô to Ga 43 (1) : 5, f. 15, TL: Jiadengyu, Altai, 1380m
Coenonympha semenovi jiadengyuica ; [MRS, 404]

Coenonympha tyderes Leech²

Tibet. See [About maps] Coenonympha tyderes ; [BOW: pl. 200, f. 5 (text only)]

16.2.2013 (39)

References:

[A. Timchenko]

Andrew Timchenko; Beijing, China (ex-Moscow, Russia)

[ALBERTA]

University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection

[About maps]

Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

[BAMONA]

Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node

[BHL]

The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!

[BMAT]

Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9

[BOW]

Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X

[BRU]

Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997; Guide to the Butterflies of Russia and adjacent territories; Hesperiidae, Papilionidae, Pieridae, Satyridae; Volume 1

[CiNii]

http://ci.nii.ac.jp/

[D. Smirnov]

Russia, Moscow; e-mail: D. Smirnov

[EBW]

Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6

[ESP]

Heppner, 1981; The dates of E. J. C. Esper's Die Schmetterlinge in Abblidungen... 1776-[1830]; Archives of Natural History (1981) 10 (2): 251-254

[Enrique Garcia-Barros]

Enrique García-Barros; Dept. Biol. (Zool.); Univ. Autonoma de Madrid

[H&R]

Higgins, L.G, Riley, N.D, 1970; A Field Guide to the Butterflies of Britain and Europe

[HBN]

Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2

[Hannu Tanner]

Hannu Tanner, Siuntio, Finland; e-mail: Hannu Tanner; http://www.kolumbus.fi/hannu.j.tanner/coleoptera/

[Jean Delacre]

http://homepage.mac.com/jdelacre

[Kimmo Silvonen]

Espoo, Finland; Kimmo's Lep Site

[Leif Wahlberg]

Julita, Sweden; e-mail: Leif Wahlberg

[MBE]

Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/

[MRS]

Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2

[Marek Michalski]

Lodz, Poland; e-mail: Marek Michalski

[Mario Maier]

Europäische Schmetterlinge; e-mail: Mario Maier

[Maurizio Elia]

Ravenna, Italy; e-mail: Maurizio Elia

[Michael Borsch]

Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes

[Michal Koupý]

Czech Republic

[Michel Tarrier]

Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org

[NAB]

Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0

[NACL]

Hodges, 1983; Check List of the Lepidoptera of America North of Mexico

[NHM]

Natural History Museum; The Global Lepidoptera Names Index

[NL4A]

Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea

[NORLEP]

Norges Sommerfugler

[Oleg Kosterin]

Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/

[Opler]

Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.

[SPRK]

Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14

[SVF]

Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)

[SZMN]

Lepidoptera; Siberian Zoological Museum

[Simon Coombes]

1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk

[Szabolcs Safian]

Hungary; Hungarian Lepidopterologial Society; Butterflies of Ghana

[Tero Piirainen]

Tampere, Finland; e-mail: Tero Piirainen

[Timo Viitanen]

http://www.kuvarkisto.com/

[USGS]

Northern Prairie Wildlife Research Center; http://www.npwrc.usgs.gov/index.htm

[Vitaly Charny]

Birmingham, Alabama, USA

[Wahlberg]

Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group

[²]

This may require parentheses or not. I don't have the necessary information for this taxon.

Some related literature:

Abadjiev, 2000

Type of Balkan butterflies. II. Eastern Large Heath, Coenonympha rhodopensis Elwes, 1900: type locality and lectotype designation Atalanta 31 (3/4): 461-466

Abadjiev, 2002

Types of Balkan butterflies in the collection of The Natural History Museum, London Neue Ent. Nachr. 23: 3-53

Alphéraky, 1881

Lépidoptères du district de Kouldjà et des montagnes environnantes. (1-3) Horae Soc. ent. Ross. 16 (3-4): 334-435, pl. 14-15 (1881), 17 (1-2): 15-103, pl. 1-3 (1882), 17 (3-4): 156-227, pl. 8-9 (1883)

Alphéraky, 1887

Diagnoses de quelques lépidoptères inédits du Thibet in Romanoff, Mem. lép. 3 : 403-406

Alphéraky, 1888

Neue Lepidopteren Stettin Ent. Ztg 49 : 66-69

Assmus, 1857

Coenonympha anaxagoras eine neue Varietät der Coenonympha iphis und ein Beitrag zur Naturgeschichte von Larentia Pyropata Hüb. Breslau Ent. Z. 10 : 9-10

O. Bang-Haas, 1927

Horae Macrolepidopterologicae Regionis Palaearcticae Horae Macrolep. Palaearct. 1: 128pp, 11pls

Blachier, 1905

Descriptions de Lépidoptères nouveaux de Maroc Bull. Soc. ent. Fr. 1905 : 212-215

Borkhausen, 1788

Naturgeschichte der europäischen Schmetterlinge nach Systematischer Ordnung. 1 Naturges. Eur. Schmett., 1: 36+288pp, 1pl

Butler, 1868

A catalogue of diurnal Lepidoptera of the family Satyridae in the collection of the British Museum Cat. diurn. Lep. Satyridae Brit. Mus. : 1-211, pl. 1-5

Butler, 1877

On Rhopalocera from Japan and Shanghai, with Descriptions of new Species Ann. Mag. nat. Hist. (4) 19 (109) : 91-97

Davenport, 1941

The Butterflies of the Satyrid Genus Coenonympha Bull. Mus. comp. Zool. Harv. 87 (4): 215-349, 10pls

dos Passos, 1935

Some butterflies of southern Newfoundland with descriptions of new subspecies Can. Ent. 67: 82-88

Westwood, [1851]

The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)

Dyar, 1903

A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723

Edwards, 1861

Descriptions of certain Species of Diurnal Lepidoptera found within the limits of the United States and of British America. - No. [1]-3 Proc. Acad. nat. Sci. Philad. 13: 160-164 (1861), 14: 54-58, 221-226 (1862)

Edwards, 1869

Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 2 : 369-376

Edwards, 1872

List of species of butterflies collected by Campbell Carrington and Wm. B. Logan, of the expedition [to Montana], in 1871. Preliminary report [5th Ann. rep.] U. S. Geological Survey of Montana, by F. V. Hayden in Hayden, Geol. Surv. Montana: 466-467

Edwards, 1881

Coenonympha elko Can. Ent. 13 (3) : 57-58

H. Edwards, 1881

On some apparently new forms of diurnal lepidoptera Papilio 1 (4) : 50-55

Elwes, 1900

On the Butterflies of Bulgaria Trans. ent. Soc. Lond. 1900 (2) : 183-206, pl. 4

Esper, 1777

Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Band 1 Die Schmett. Th. I, Bd. 1 (1): 1-40, pl. 1-6 (1776), (2): 41-72, pl. 7-12 (1776), (3): 73-104, pl. 13-18 (1777), (4): 105-120, pl. 19-24 (1777), (5): 121-144, pl. 25-30 (1777), (6): 145-176, pl. 31-36 (1777), (7): 177-200, pl. 37-46 (1778), (8): 201-216, pl. 43-48 (1778), (9): 217-388, pl. 49-50 (1779)

Esper, 1782; Esper, 1784

Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Fortsetzung. Band 2 Die Schmett. Th. I, Bd. 2 (1): 1-36, pl. 51-56 (1780), (2): 37-68, pl. 57-62 (1781), (3): 69-100, pl. 63-68 (1781), (4): 101-124, pl. 69-74 (1781), (5): 125-140, pl. 75-80 (1782), (7): 141-156, pl. 81-82 (1783), (8): 157-172, pl. 83-88 (1783), (9): 173-184, pl. 89-92 (1784), (28): 185-190, pl. 93 (1786)

Esper, 1794

Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Supplement Theil 1. Abschnitt 1 Die Schmett., Suppl. Th 1 (1-2): 1-8, pl. 94 (1789), (3-4): 9-60, pl. 95-101 (1789), (5-6): 61-68, pl. 102-106 (1794), ([7]): [69-88], pl. [107-109] (1798), (8-9): 89-104, pl. 110-112 (1800), (10): 105-120, pl. 113-116 ([1803-1804])

Esper, 1805

Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Supplement Theil 2 Die Schmett., Th I, Suppl. Th. 2 (11): 1-24, pl. 117-122 (1805), (12): 25-48, pl. 123-126 [1830]

Erschoff, 1874

Travels in Turkestan. Volume 2. Zoogeographical Investigations. Lepidoptera [in Russian] in Fedschenko, Travels in Turkestan. 2 (5) : 1-128, pl. 1-6

Eversmann, 1843

Quaedam lepidopterorum species novae in montibus Uralensibus et Altaicus habitantes nunc descriptae et depictae Bull. Soc. Imp. Nat. Moscou 16 (3) : 535-553, 4pls

Eversmann, 1851

Description de quelques nouvelles espéce de Lépidoptéres de la Russie Bull. Soc. imp. Nat. Moscou 24 (2) : 610-644

Fabricius, 1777

Genera Insectorum eorumque characters naturales secundum numerum, figuram, situm et proportionem ... Genera Insectorum : 1-310

Freyer, 1845

Neuere Beiträge zur Schmetterlingskunde mit Abbildungen nach der Natur. (65-80) Neuere Beitr. Schmett. 5 : 1-40 (1842), : 41-92 (1844), : 93-164 (1845)

Heyne, [1894]

Satyridae, Lycaenidae in Rühl, Die palaearktischen Grossschmetterlinge und ihre Naturgeschichte 1 (Tagfalter): 385-857

Holland, 1900

Alaska Insects Ent. News 11 (3): 381-389 (part I), (4): 416-423 (part II)

Huang & Murayama, 1992

Butterflies of Xinjiang province, China Tyô to Ga 43 (1) : 1-22

Hübner, 1804

Sammlung europäischer Schmetterlinge. I. Papiliones - Falter ("Erste Band") Samml. eur. Schmett. [1] : 3-74, : pl. 1-88, f. 1-446 (1799-1800), : pl. 89-96, f. 447-490 (1800-1803), : pl. 97-114, f. 491-588 (1803-1804), : pl. 115-119, f. 589-612 (1805), : pl. 120-124, f. 613-636 (1805-1806), : pl. 125-128, f. 637-652 (1806-1808), : pl. 129-144, f. 653-727 (1808-1813), : pl. 145-147, f. 728-741 (1814-1816), : pl. 148-150, f. 742-756 (1816-1817), : pl. 151, f. 757-762 (1818-1819), : pl. 152-154, f. 757-774 (1819-1822), : pl. 155, f. 775-778 (1822-1823), : pl. 156-161, f. 779-806 (1823), : pl. 162-175, f. 807-874 (1823-1824), : pl. 176, f. 869-874 (1824-1825), : pl. 177-181, f. 875-895 (1825-1826), : pl. 182-187, f. 896-981 (1827-1828), : pl. 188-195, f. 922-968 (1828-1832), : pl. 196, f. 969-974 (1832-1833), : pl. 197-201, f. 975-997 (1834-1836), : pl. 202-203, f. 998-1009 (1836-1838), : pl. 204-207, f. 1010-1029 (1837-1838)

Hübner, [1819]

Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])

Krulikowsky, 1908

Einige neue Varietäten und Aberrationen der Lepidopteren des östlichen Russlands Soc. Ent. 23 (1): 2-3, (2): 11-12, (3): 18

Lederer, 1870

Contributions a la faune des Lepidopteres de la Transcaucasie Ann. Soc. ent. Belg. 13 : 17-54, pl. 1-2

Leech, [1892]

Butterflies from China, Japan, and Corea Butts China Japan Corea (1): 1-297, pl. 1-28 (1892-1894), (2): 297-681, pl. 29-43 (1893-1894)

Linnaeus, 1758

Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824

Matsumura, 1927

A list of the butterflies of Corea, with description of new species, subspecies and aberrations Insecta Matsumurana 1 (4) : 159-170

McDunnough, 1939

A new Coenonympha race from northeastern New Brunswick Can. ent. 71: 266

Müller, 1764

Fauna Insectorum Fridrichsdalina sive Methodica Descriptio Insectorum Agri Fridrichsdalensis cum Characteribus Genericis et Specificis, Nominibus Trivialibus, Locis Natalibus, Iconibus Allegatis, Novsque Pluribus Speciebus Additis Fauna Ins. Fridr.: 96pp

Oberthür, 1874

Lépidoptères nouveaux d'Algérie. 1. Diurnes Petites Nouv. Ent. 1: 412-413

Oberthür, 1910

Notes pour servir à établir la Faune Française et Algérienne des Lépidoptères Ètud. Lépid. Comp. 4 : 15-691, pl. 35-58

Oberthür, 1922

Les Lépidoptères du Maroc Étud. Lépid. Comp. 19 (1): 13-402, (2): pl. 530-546

Pierret, 1837

Description de deux nouvelles espèces du genre Satyre (Latr.) Ann. Soc. ent. Fr. 6 : 303-307, pl. 12

Poda, 1761

Insecta Musei Graecensis, quae in ordines, genera et species juxta Systema Naturae Caroli Linnaei digessit Ins. Mus. Graecensis: 8+127, 2pls

Porter & Mattoon, 1989

A new subspecies of Coenonympha tullia (Müller) confined to the coastal dunes of northern California J. Lep. Soc. 43 (3): 229-238

de Prunner, 1798

Lepidoptera Pedemontana illustrata. Turin. 1 Lepid. Pedemont.: 52+124

Reakirt, 1866

Coloradian butteflies Proc. Ent. Soc. Philad. 6 (1) : 122-151

Rothschild, 1917

Supplemental notes to Mr. Charles Oberthür's Faune des Lépidoptères de la Barbarie, with lists of the specimens contained in the Tring Museum Novit. Zool. 24 (1): 61-120, : 325-373, : 393-409

Rothschild, 1925

Critical List of the collection of Algerian Lepidoptera of the late Captain N. J. E. Holl. Novit. Zool. 32 : 195-229

Rottemburg, 1775

Unmertungen zu den Hufnagelifchen Tabellen der Schmetterlinge Der Naturforscher 6 : 1-34

Seitz, 1908

Die Großschmetterlinge des Palaearktischen Faunengebietes. 1. Die Palaearktischen Tagfalter Gross-Schmett. Erde 1 : 1-379

Sheljuzhko, 1929

Einige neue palaearktische Lepidopteren-Formen Mitt. Münch. Ent. Ges. 19 : 347-362, pl. 27-28

Staudinger, 1870

Beschreibung neuer Lepidopteren des europäischen faunegebiets Berl. Ent. Z. 14 : 97-132, 193-208, (3/4): 273-330 (1871)

Staudinger, 1886

Centralasiatische Lepidopteren Stett. Ent. Ztg. 47 (4-6): 193-215, (7-9): 225-256

Stoll, [1782]

Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)

Thunberg, 1791

Dissertio Entomologica sistens Insecta Suecica, ... Diss. Ent. sistens Insecta Suecica (1): 1-24 (1 plate) (1784), (2): [25-31] ??], 32-46 (9 unnumbered) (1791), (3): [47],48-52 (2 unnumbered) (1792), (4): 53-62 (1792)

Tutt, 1904

Coenonympha mathewi, n.sp. Ent. Rec. J. Var. 16 : 308-309

Verity, 1916

The Brish Races of Butterflies: their relationships and nomeclature Ent. Rec. J. Var. 28 (5): 97-102, (6): 128-133, (7-8): 165-174

Webster, 1998

The life history of the Maritime Ringlet, Coenonympha tullia nipisquit J. Lep. Soc. 52 (4): 345-355

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.