[BRU2]; Tuzov, Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000
Guide to the Butterflies of Russia and adjacent territories: Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae
Butts. Russia adj. terr. 2
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