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Carcharodus Hübner, [1819]
Carcharodus Hübner, [1819]; Verz. bek. Schmett. (7): 110, TS: Papilio alceae Esper
Reverdinus Ragusa, 1919; Naturalista Siciliano 23 (7/12): 172, TS: Papilio altheae Hübner
Lavatheria Verity, 1940; Le Farfalle diurn. d'Italia 1: 11, 22, TS: Papilio lavatherae Esper
Carcharodus (Pyrginae) ; [AfrL]

Carcharodus alceae (Esper, 1780)
SEU, CEU, Temperate Asia (S.Siberia - Arabia, N.India). See [About maps] Mallow Skipper Piquitos castana Warcabnik ślazowiec
Papilio alceae Esper, 1780; Die Schmett. Th. I, Bd. 2 (1): 4, pl. 51, f. 3, TL: Erlangen, Germany
malvarum Hoffmannsegg, 1804
Carcharodus alceae, Moths and Butterflies of Europe [MBE]
Carcharodus alceae, Extracts from [BRU] by [Ilya Osipov]
Carcharodus alceae, Captain's European Butterfly Guide [Simon Coombes]
Carcharodus alceae ; [BOW: pl. 11, f. 11]; [BRU: 108, pl. 2, f. 1-3]; [BMAT: 106 (all N.African records are Carcharodus tripolina)]; [AfrL]; [Otakar Kudrna]
[alceae-1.jpg] 600x476 (~104Kb) Italy, Ravenna, 11 May 2006, Photo © Maurizio Elia [alceae.jpg] 500x324 (~47Kb) Turunc, SW Turkey 14/04/95, Photo © Simon Coombes

Larva on Malva, Althaea, Lavatera, Hibiscus, (etc) [BRU] (in Spain) Malva rosea [BMAT]


C. a. wissmanni Warnecke, 1934
Carcharodus alceae wissmanni ; [AfrL]
C. a. swinhoei Watson, 1893
Afghanistan, NW.India, ... Carcharodus swinhoei Watson, 1893; Proc. Zool. Soc. Lond. 1893 (1) : 68

Carcharodus tripolina (Verity, 1925)
SWEU, N.Africa. See [About maps] Erynnis alceae tripolina Verity, 1925; Ent. Rec. J. Var. 37 : 54, TL: Garian plateau (Libya)
Carcharodus tripolina ; [BMAT: 94, pl. 31, f. 6-10]

Larva on Malva sylvestris, (in Egypt) Althaea sp. [BMAT]


Carcharodus lavatherae (Esper, 1783)
NAF, SEU, Asia Minor - Iran. See [About maps] Marbled Skipper Ziestfalter Piquitos clara
Papilio lavatherae Esper, 1783; Die Schmett. Th. I, Bd. 2 (7): 148, pl. 82, f. 4, TL: France & Switzerland
Carcharodus lavatherae, Moths and Butterflies of Europe [MBE]
Carcharodus lavatherae, Extracts from [BRU] by [Ilya Osipov]
Carcharodus lavatherae, Captain's European Butterfly Guide [Simon Coombes]
Carcharodus lavatherae ; [BOW: pl. 11, f. 11 (name)]; [BRU: 108, pl. 2, f. 4-6]; [Otakar Kudrna]

Larva on Stachys [H&R], Stachys recta [BRU]

C. l. rufescens Oberthür, 1915
Morocco (Middle Atlas), Algeria, Tunisia? Carcharodus lavaterae[sic] var. rufescens Oberthür, 1915; Étud. Lépid. Comp. 10 : 407, pl. 296, f. 4433, TL: Sebdou (Algeria)
Carcharodus lavaterae[sic] internicus Rothschild, 1914; Novit. Zool. 21: 311, TL: Aïn Sefra, Guelt-es-Stel (Algeria)
Erynnis lavatherae australissima Verity, 1925; Ent. Rec. J. Var. 37 : 41, TL: Sebdou, Lambessa (Algeria)
Carcharodus lavatherae rufescens ; [BMAT: 95, pl. 31, f. 11-15]

Carcharodus baeticus (Rambur, 1840)
SWEU, it, Asia Minor, Iran. See [About maps] Southern Marbled Skipper Piquitos
marrubii Rambur, 1840
boeticus[sic!] ;
Carcharodus baeticus ; [Otakar Kudrna]

Larva on Marrubium vulgare, Ballota foetida, (etc) [H&R]


Subgenus Reverdinus

Carcharodus floccifera (Zeller, 1847)
SEU, Bulgaria, S.Siberia, Asia Minor. See [About maps] Tufted Skipper Piquitos serrana Warcabnik szantawiec
Hesperia floccifera Zeller, 1847; Isis Jena, 1847 (1): 286, TL: Syracuse, Sicily
Papilio altheae Hübner, [1800-1803]; Samml. eur. Schmett. [1] : pl. 90, f. 452-453 (preocc. Papilio altheae Esper, [1783])
Carcharodus imperator Hemming, 1934; Stylops 3 : 99 (repl. Papilio altheae Hübner, [1800-1803])
alchymillae Hemming, 1936
gemina Lederer, 1852
Carcharodus floccifera, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Carcharodus flocciferus, Moths and Butterflies of Europe [MBE]
Carcharodus flocciferus, Extracts from [BRU] by [Ilya Osipov]
Carcharodus flocciferus, Captain's European Butterfly Guide [Simon Coombes]
Carcharodus flocciferus ; [BRU: 109, pl. 2, f. 7-9]; [Otakar Kudrna]
[floccifera-1u.jpg] 533x400 (~40Kb) upperside An open pine forest on a high left bank of the Berd' River 1 km upstream of the village Starososedovo. Iskitim district, Novosibirsk Province, West Siberia, Russia. 2nd July 1994, Photo © Oleg Kosterin

Larva on Marrubium peregrinum, M. vulgare, Stachys officinalis, Thymus roegneri [BRU], Stachys alpina, S. recta [BMAT]


C. f. habiba (Kauffmann, 1955)
Morocco (W.Rif Mts) Reverdinus floccifer habiba Kauffmann, 1955; Mitt. schweiz. ent. Ges. 28 (3): 288, TL: Bab el Taza (Morocco)
Carcharodus floccifera habiba ; [BMAT: 95, pl. 31, f. 16-18]

Carcharodus orientalis Reverdin, 1913
Montenegro, Greece, Asia Minor, Palestine, N.Iran. See [About maps] Oriental Skipper
Carcharodus orientalis Reverdin, 1913; Bull. Soc. Lép. Genève, 2 (4): 230, f. 4, pl. 21, f. 14, TL: Peloponnesos, Greece
Carcharodus orientalis, Moths and Butterflies of Europe [MBE]
Carcharodus orientalis, Extracts from [BRU] by [Ilya Osipov]
Carcharodus orientalis ; [BRU, 109]; [Otakar Kudrna]
C. o. teberdininus Devyatkin, 1990
Caucasus Major

Carcharodus stauderi Reverdin, 1913
Morocco - Cyrenaica, Greece, Asia Minor - Palestine, N.Iran. See [About maps] Carcharodus stauderi Reverdin, 1913; Bull. Soc. Lép. Genève, 2 (4): 225, pl. 21, f. 5, 12, TL: El Kantara (Algeria)
Carcharodus stauderi, Extracts from [BRU] by [Ilya Osipov]
Carcharodus stauderi ; [BRU: 109, pl. 2, f. 13-15]; [Otakar Kudrna]

Larva on Marrubium vulgare, Ballota foetida, (etc) [H&R] Nepeta floccosa [BRU]

N.Africa Erynnis stauderi f. fulvissima Verity, 1925; Ent. Rec. J. Var. 37 : 43, TL: Algeria
Erynnis stauderi f. obscura Verity, 1925; Ent. Rec. J. Var. 37 : 44, TL: Algeria
Carcharodus stauderi romeii Rothschild, 1933; Novit. Zool. 38 : 323, TL: Ketama (Morocco)
Reverdinus stauderi f. rungsi Picard, 1950; Bull. Soc. Sci. nat. Maroc 28: 118-119, TL: Ifrane (Morocco)
Carcharodus stauderi stauderi ; [BMAT: 95, pl. 31, f. 19-20]

Larva on Marrubium vulgare, (in Hoggar) Marrubium deserti?, Ballota hirsuta, (in Sinai) Phlomis aurea [BMAT]

C. s. ambigua Verity, 1925
S.Transcaucasia

Carcharodus dravira (Moore, [1875])
N.Iran - Pakistan, N.India. See [About maps] Pyrgus dravira Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4) : 576, pl. 67, f. 5, TL: Kashmir
balucha Evans, 1932
Carcharodus dentatus Alberti, 1940; Mitt. Münch. ent. Ges. 30 : 236, pl. 2, f. 5d, TL: Badachschan, Sebaktal, Alpenwiesenzone, 2800-3000m
Carcharodus dravira, Extracts from [BRU] by [Ilya Osipov]
Pyrgus dravira ; de Nicéville, [1884], J. Asiat. Soc. Bengal 52 Pt.II (2/4) : 88, pl. 10, f. 5 ♀
Carcharodus dravira ; [BRU: 110, pl. 2, f. 16-18]


7.10.2011 (8)

References:
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU]
Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997; Guide to the Butterflies of Russia and adjacent territories; Hesperiidae, Papilionidae, Pieridae, Satyridae; Volume 1
[ESP]
Heppner, 1981; The dates of E. J. C. Esper's Die Schmetterlinge in Abblidungen... 1776-[1830]; Archives of Natural History (1981) 10 (2): 251-254
[H&R]
Higgins, L.G, Riley, N.D, 1970; A Field Guide to the Butterflies of Britain and Europe
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Ilya Osipov]
The insects from former USSR for collectors Web site; e-mail: Ilya Osipov; http://www.osipov.org/insects/
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[Maurizio Elia]
Ravenna, Italy; e-mail: Maurizio Elia
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk


Some related literature:
Alberti, 1940
Über einige neue oder wenig bekannte palaearktische Hesperiiden nebst Revision der Hesperia staudingeri-Gruppe Mitt. Münch. ent. Ges. 30 : 235-254, 10fgs
de Nicéville, [1884]
On new and little known Rhopalocera from the Indian Region J. Asiat. Soc. Bengal 52 Pt.II (2/4) : 65-91, pl. 1, 9-10
Esper, 1780; Esper, 1783
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Fortsetzung. Band 2 Die Schmett. Th. I, Bd. 2 (1): 1-36, pl. 51-56 (1780), (2): 37-68, pl. 57-62 (1781), (3): 69-100, pl. 63-68 (1781), (4): 101-124, pl. 69-74 (1781), (5): 125-140, pl. 75-80 (1782), (7): 141-156, pl. 81-82 (1783), (8): 157-172, pl. 83-88 (1783), (9): 173-184, pl. 89-92 (1784), (28): 185-190, pl. 93 (1786)
Hemming, 1934
Some notes in the nomenclature of Palaearctic and African Rhopalocera Stylops 3 : 97-99
Hübner, [1800-1803]
Sammlung europäischer Schmetterlinge. I. Papiliones - Falter ("Erste Band") Samml. eur. Schmett. [1] : 3-74, : pl. 1-88, f. 1-446 (1799-1800), : pl. 89-96, f. 447-490 (1800-1803), : pl. 97-114, f. 491-588 (1803-1804), : pl. 115-119, f. 589-612 (1805), : pl. 120-124, f. 613-636 (1805-1806), : pl. 125-128, f. 637-652 (1806-1808), : pl. 129-144, f. 653-727 (1808-1813), : pl. 145-147, f. 728-741 (1814-1816), : pl. 148-150, f. 742-756 (1816-1817), : pl. 151, f. 757-762 (1818-1819), : pl. 152-154, f. 757-774 (1819-1822), : pl. 155, f. 775-778 (1822-1823), : pl. 156-161, f. 779-806 (1823), : pl. 162-175, f. 807-874 (1823-1824), : pl. 176, f. 869-874 (1824-1825), : pl. 177-181, f. 875-895 (1825-1826), : pl. 182-187, f. 896-981 (1827-1828), : pl. 188-195, f. 922-968 (1828-1832), : pl. 196, f. 969-974 (1832-1833), : pl. 197-201, f. 975-997 (1834-1836), : pl. 202-203, f. 998-1009 (1836-1838), : pl. 204-207, f. 1010-1029 (1837-1838)
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Moore, [1875]
Descriptions of new Asiatic Lepidoptera Proc. zool. Soc. Lond. 1874 (4) : 565-579, pl. 66-67
Oberthür, 1915
Faune des Lépidoptères de la Barbarie, Algerie Étud. Lépid. Comp. 10 : 1-459, pl. 276-277,277bis,278-315
Reverdin, 1913
Notes sur les genres Hesperia et Carcharodus Bull. Soc. Lép. Genève, 2 (4): 212-234, pl. 21-22
Rothschild, 1933
On a collection of Lepidoptera from Spanish Morocco Novit. Zool. 38 : 315-330
Verity, 1925
New races and forms of Palaearctic Grypocera Ent. Rec. J. Var. 37 : 41-44,54-57,72-76
Watson, 1893
A proposed classification of the Hesperiidae, with a revision of the genera Proc. Zool. Soc. Lond. 1893 (1) : 3-133, pl. 1-3


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.